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Note 1 - Summary of Revenues (Details) - USD ($) $ in Thousands | 3 Months Ended
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Mar. 31, 2016 | Mar. 31, 2015
US Department of Health and Human Services [Member] |  |
Collaborative and other research and development revenues: |  |
Collaborative and other research and development revenues | $ 2,324 | $ 4,475
Shionogi and Co. Ltd [Member] |  |
Collaborative and other research and development revenues: |  |
Collaborative and other research and development revenues | 296 | $ 296
Seqirus UK Limited [Member] |  |
Collaborative and other research and development revenues: |  |
Collaborative and other research and development revenues | $ 310 |
Product sales, net |  | $ 537
Royalty revenue | $ 1,890 | 1,518
Collaborative and other research and development revenues | 2,930 | 4,771
Revenues | $ 4,820 | $ 6,826

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