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Consolidated Statements of Comprehensive Loss (Unaudited) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended
	Mar. 31, 2017	Mar. 31, 2016
Revenues
Royalty revenue	$ 6,321	$ 1,890
Collaborative and other research and development	3,116	2,930
Total revenues	9,437	4,820
Expenses
Research and development	16,770	20,579
General and administrative	3,058	3,212
Royalty	294	77
Total operating expenses	20,122	23,868
Loss from operations	(10,685)	(19,048)
Interest and other income	109	439
Interest expense	(2,100)	(1,470)
Loss on foreign currency derivative	(1,543)	(2,753)
Net loss	$ (14,219)	$ (22,832)
Basic and diluted net loss per common share (in dollars per share)	$ (0.19)	$ (0.31)
Weighted average shares outstanding (in shares)	75,167	73,601
Unrealized gain on available for sale investments	$ 9	$ 254
Comprehensive loss	$ (14,210)	$ (22,578)

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Amount after tax of increase (decrease) in equity from transactions and other events and circumstances from net income and other comprehensive income, attributable to parent entity. Excludes changes in equity resulting from investments by owners and distributions to owners.

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The amount of net income or loss for the period per each share in instances when basic and diluted earnings per share are the same amount and reported as a single line item on the face of the financial statements. Basic earnings per share is the amount of net income or loss for the period per each share of common stock or unit outstanding during the reporting period. Diluted earnings per share includes the amount of net income or loss for the period available to each share of common stock or common unit outstanding during the reporting period and to each share or unit that would have been outstanding assuming the issuance of common shares or units for all dilutive potential common shares or units outstanding during the reporting period.

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The aggregate costs incurred (1) in a planned search or critical investigation aimed at discovery of new knowledge with the hope that such knowledge will be useful in developing a new product or service, a new process or technique, or in bringing about a significant improvement to an existing product or process; or (2) to translate research findings or other knowledge into a plan or design for a new product or process or for a significant improvement to an existing product or process whether intended for sale or the entity's use, during the reporting period charged to research and development projects, including the costs of developing computer software up to the point in time of achieving technological feasibility, and costs allocated in accounting for a business combination to in-process projects deemed to have no alternative future use.

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