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Consolidated Statements of Comprehensive Loss (Unaudited) - USD ($) shares in Thousands, $ in Thousands | 3 Months Ended | 6 Months Ended
--- | --- | ---
Jun. 30, 2017 | Jun. 30, 2016 | Jun. 30, 2017 | Jun. 30, 2016
Revenues |  |  |  |
Royalty revenue | $ 489 | $ 629 | $ 6,810 | $ 2,519
Collaborative and other research and development | 2,610 | 4,158 | 5,726 | 7,088
Total revenues | 3,099 | 4,787 | 12,536 | 9,607
Expenses |  |  |  |
Research and development | 15,759 | 14,166 | 32,529 | 34,745
General and administrative | 2,834 | 2,724 | 5,892 | 5,936
Royalty | 22 | 27 | 316 | 104
Total operating expenses | 18,615 | 16,917 | 38,737 | 40,785
Loss from operations | (15,516) | (12,130) | (26,201) | (31,178)
Interest and other income | 203 | 147 | 312 | 586
Interest expense | (2,094) | (1,421) | (4,194) | (2,891)
Gain (loss) on foreign currency derivative | 521 | (2,877) | (1,022) | (5,630)
Net loss | $ (16,886) | $ (16,281) | $ (31,105) | $ (39,113)
Basic and diluted net loss per common share (in dollars per share) | $ (0.21) | $ (0.22) | $ (0.40) | $ (0.53)
Weighted average shares outstanding (in shares) | 80,418 | 73,695 | 77,807 | 73,648
Unrealized loss on available for sale investments | $ (12) | $ (2) | $ (3) | $ 252
Comprehensive loss | $ (16,898) | $ (16,283) | $ (31,108) | $ (38,861)

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- Definition Amount after tax of increase (decrease) in equity from transactions and other events and circumstances from net income and other comprehensive income, attributable to parent entity. Excludes changes in equity resulting from investments by owners and distributions to owners. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 220 -SubTopic 10 -Section 45 -Paragraph 5 -URI http://asc.fasb.org/extlink&oid=84228862&loc=d3e557-108580 + Details
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