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Note 1 - Significant Accounting Policies - Summary of Revenues (Details) - USD ($) $ in Thousands | 3 Months Ended | 6 Months Ended
--- | --- | ---
Jun. 30, 2017 | Jun. 30, 2016 | Jun. 30, 2017 | Jun. 30, 2016
Royalty revenue | $ 489 | $ 629 | $ 6,810 | $ 2,519
Collaborative and other research and development revenues | 2,610 | 4,158 | 5,726 | 7,088
Revenues | 3,099 | 4,787 | 12,536 | 9,607
US Department of Health and Human Services [Member] |  |  |  |
Collaborative and other research and development revenues | 2,161 | 3,709 | 2,815 | 6,033
Shionogi and Co. Ltd [Member] |  |  |  |
Collaborative and other research and development revenues | 296 | 296 | 592 | 592
Seqirus UK Limited [Member] |  |  |  |
Collaborative and other research and development revenues | $ 153 | $ 153 | $ 2,319 | $ 463

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