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Consolidated Statements of Comprehensive Loss (Unaudited) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2017	Jun. 30, 2016	Jun. 30, 2017	Jun. 30, 2016
Revenues
Royalty revenue	$ 489	$ 629	$ 6,810	$ 2,519
Collaborative and other research and development	2,610	4,158	5,726	7,088
Total revenues	3,099	4,787	12,536	9,607
Expenses
Research and development	15,759	14,166	32,529	34,745
General and administrative	2,834	2,724	5,892	5,936
Royalty	22	27	316	104
Total operating expenses	18,615	16,917	38,737	40,785
Loss from operations	(15,516)	(12,130)	(26,201)	(31,178)
Interest and other income	203	147	312	586
Interest expense	(2,094)	(1,421)	(4,194)	(2,891)
Gain (loss) on foreign currency derivative	521	(2,877)	(1,022)	(5,630)
Net loss	$ (16,886)	$ (16,281)	$ (31,105)	$ (39,113)
Basic and diluted net loss per common share (in dollars per share)	$ (0.21)	$ (0.22)	$ (0.40)	$ (0.53)
Weighted average shares outstanding (in shares)	80,418	73,695	77,807	73,648
Unrealized loss on available for sale investments	$ (12)	$ (2)	$ (3)	$ 252
Comprehensive loss	$ (16,898)	$ (16,283)	$ (31,108)	$ (38,861)

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