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Note 1 - Significant Accounting Policies - Cumulative Effective Adopted Accounting Pronouncement (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended |  |
--- | --- | --- | ---
Mar. 31, 2018 | Mar. 31, 2017 | Jan. 01, 2018 | Dec. 31, 2017
Deferred collaboration expense | $ 152 |  | $ 152 | $ 210
Deferred revenue | 7,300 |  | 7,300 | 8,484
Accumulated deficit | (656,494) |  | (630,717) | $ (631,843)
Revenues | 3,976 | $ 9,437 |  |
Research and development | 18,441 | 16,770 |  |
Net loss | $ (25,777) | $ (14,219) |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.26) | $ (0.19) |  |
Accounting Standards Update 2014-09 [Member] |  |  |  |
Deferred collaboration expense |  |  | (58) |
Deferred revenue |  |  | (1,184) |
Accumulated deficit |  |  | $ 1,126 |
Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |
Deferred collaboration expense | $ 44 |  |  |
Deferred revenue | 888 |  |  |
Accumulated deficit | 282 |  |  |
Research and development | 14 |  |  |
Net loss | $ 282 |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ 0 |  |  |
Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |
Deferred collaboration expense | $ 196 |  |  |
Deferred revenue | 8,188 |  |  |
Accumulated deficit | (656,212) |  |  |
Research and development | 18,455 |  |  |
Net loss | $ (25,495) |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.26) |  |  |
Collaborative and Other Research and Development [Member] |  |  |  |
Revenues | $ 315 | $ 3,116 |  |
Collaborative and Other Research and Development [Member] | Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |
Revenues | 296 |  |  |
Collaborative and Other Research and Development [Member] | Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |
Revenues | $ 611 |  |  |

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