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Note 1 - Significant Accounting Policies - Cumulative Effective Adopted Accounting Pronouncement (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 6 Months Ended |  |
--- | --- | --- | --- | ---
Jun. 30, 2018 | Jun. 30, 2017 | Jun. 30, 2018 | Jun. 30, 2017 | Jan. 01, 2018 | Dec. 31, 2017
Deferred collaboration expense | $ 12 |  | $ 12 |  | $ 152 | $ 210
Deferred revenue | 300 |  | 300 |  | 7,300 | 8,484
Accumulated deficit | (674,940) |  | (674,940) |  | (630,717) | $ (631,843)
Revenues | 12,494 | $ 3,099 | 16,470 | $ 12,536 |  |
Research and development | 21,010 | 15,759 | 39,451 | 32,529 |  |
Net loss | $ (18,446) | $ (16,886) | $ (44,223) | $ (31,105) |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.19) | $ (0.21) | $ (0.45) | $ (0.40) |  |
Accounting Standards Update 2014-09 [Member] |  |  |  |  |  |
Deferred collaboration expense |  |  |  |  | (58) |
Deferred revenue |  |  |  |  | (1,184) |
Accumulated deficit |  |  |  |  | $ 1,126 |
Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |  |  |
Deferred collaboration expense | $ 29 |  | $ 29 |  |  |
Deferred revenue | 592 |  | 592 |  |  |
Accumulated deficit | 563 |  | 563 |  |  |
Research and development | 14 |  | 29 |  |  |
Net loss | $ 282 |  | $ 563 |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ 0.01 |  | $ 0.01 |  |  |
Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |  |  |
Deferred collaboration expense | $ 41 |  | $ 41 |  |  |
Deferred revenue | 892 |  | 892 |  |  |
Accumulated deficit | (674,377) |  | (674,377) |  |  |
Research and development | 21,024 |  | 39,480 |  |  |
Net loss | $ (18,164) |  | $ (43,660) |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.18) |  | $ (0.44) |  |  |
Collaborative and Other Research and Development [Member] |  |  |  |  |  |
Revenues | $ 12,352 | $ 2,610 | $ 12,667 | $ 5,726 |  |
Collaborative and Other Research and Development [Member] | Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |  |  |
Revenues | 296 |  | 592 |  |  |
Collaborative and Other Research and Development [Member] | Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |  |  |
Revenues | $ 12,648 |  | $ 13,259 |  |  |

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