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Note 1 - Significant Accounting Policies - Cumulative Effective Adopted Accounting Pronouncement (Details) - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 9 Months Ended |  |
--- | --- | --- | --- | ---
Sep. 30, 2018 | Sep. 30, 2017 | Sep. 30, 2018 | Sep. 30, 2017 | Jan. 01, 2018 | Dec. 31, 2017
Deferred collaboration expense | $ 8 |  | $ 8 |  | $ 152 | $ 210
Deferred revenue | 200 |  | 200 |  | 7,300 | 8,484
Accumulated deficit | (704,537) |  | (704,537) |  | (630,717) | $ (631,843)
Revenues | 1,454 | $ 8,760 | 17,924 | $ 21,296 |  |
Research and development | 22,006 | 17,509 | 61,457 | 50,038 |  |
Net loss | $ (29,597) | $ (15,134) | $ (73,820) | $ (46,239) |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.28) | $ (0.18) | $ (0.73) | $ (0.58) |  |
Accounting Standards Update 2014-09 [Member] |  |  |  |  |  |
Deferred collaboration expense |  |  |  |  | (58) |
Deferred revenue |  |  |  |  | (1,184) |
Accumulated deficit |  |  |  |  | $ 1,126 |
Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |  |  |
Deferred collaboration expense | $ 19 |  | $ 19 |  |  |
Deferred revenue | 396 |  | 396 |  |  |
Accumulated deficit | 378 |  | 378 |  |  |
Research and development | 14 |  | 44 |  |  |
Net loss | $ 282 |  | $ 846 |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ 0 |  | $ 0.01 |  |  |
Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |  |  |
Deferred collaboration expense | $ 27 |  | $ 27 |  |  |
Deferred revenue | 596 |  | 596 |  |  |
Accumulated deficit | (704,159) |  | (704,159) |  |  |
Research and development | 22,021 |  | 61,501 |  |  |
Net loss | $ (29,315) |  | $ (72,974) |  |  |
Basic and diluted net loss per common share (in dollars per share) | $ (0.28) |  | $ (0.72) |  |  |
Collaborative and Other Research and Development [Member] |  |  |  |  |  |
Revenues | $ 931 | $ 6,817 | $ 13,598 | $ 12,543 |  |
Collaborative and Other Research and Development [Member] | Difference between Revenue Guidance in Effect before and after Topic 606 [Member] |  |  |  |  |  |
Revenues | 296 |  | 888 |  |  |
Collaborative and Other Research and Development [Member] | Calculated under Revenue Guidance in Effect before Topic 606 [Member] |  |  |  |  |  |
Revenues | $ 1,227 |  | $ 14,486 |  |  |

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