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Note 1 - Significant Accounting Policies and Concentrations of Risk - Cumulative Effective Adopted Accounting Pronouncement (Details) - USD ($) $ / shares in Units, $ in Thousands	3 Months Ended								12 Months Ended
	Dec. 31, 2018	Sep. 30, 2018	Jun. 30, 2018	Mar. 31, 2018	Dec. 31, 2017	Sep. 30, 2017	Jun. 30, 2017	Mar. 31, 2017	Dec. 31, 2018	Dec. 31, 2017	Dec. 31, 2016	Jan. 01, 2018
Deferred collaboration expense	$ 9				$ 210				$ 9	$ 210		$ 152
Deferred revenue	221				8,484				221	8,484		7,300
Accumulated deficit	(731,969)				(631,843)				(731,969)	(631,843)		(630,717)
Revenues	2,729	$ 1,454	$ 12,494	$ 3,976	3,890	$ 8,760	$ 3,099	$ 9,437	20,653	25,186	$ 26,353
Research and development									84,888	66,962	61,008
Net loss	$ (27,432)	$ (29,597)	$ (18,446)	$ (25,777)	$ (19,543)	$ (15,134)	$ (16,886)	$ (14,219)	$ (101,252)	$ (65,782)	$ (55,144)
Basic and diluted net loss per common share (in dollars per share)	$ (0.25)	$ (0.28)	$ (0.19)	$ (0.26)	$ (0.20)	$ (0.18)	$ (0.21)	$ (0.19)	$ (0.98)	$ (0.78)	$ (0.75)
Accounting Standards Update 2014-09 [Member]
Deferred collaboration expense												(58)
Deferred revenue												(1,184)
Accumulated deficit												$ 1,126
Difference between Revenue Guidance in Effect before and after Topic 606 [Member]
Deferred collaboration expense
Deferred revenue
Accumulated deficit
Research and development									58
Net loss									$ 1,126
Basic and diluted net loss per common share (in dollars per share)									$ 0.01
Calculated under Revenue Guidance in Effect before Topic 606 [Member]
Deferred collaboration expense	9								$ 9
Deferred revenue	221								221
Accumulated deficit	$ (731,969)								(731,969)
Research and development									84,946
Net loss									$ (100,126)
Basic and diluted net loss per common share (in dollars per share)									$ (0.97)
Collaborative and Other Research and Development [Member]
Revenues									$ 14,552	$ 13,142	$ 14,402
Collaborative and Other Research and Development [Member] | Difference between Revenue Guidance in Effect before and after Topic 606 [Member]
Revenues									1,184
Collaborative and Other Research and Development [Member] | Calculated under Revenue Guidance in Effect before Topic 606 [Member]
Revenues									$ 15,736

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