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Condensed Consolidated Statements of Stockholders' Deficit - USD ($) shares in Thousands, $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Other Comprehensive Income | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Common stock, shares outstanding, beginning balance (in shares) at Dec. 31, 2022 |  | 187,906 |  |  |
Beginning balance at Dec. 31, 2022 | $ (294,597) | $ 1,879 | $ 1,158,118 | $ 26 | $ (1,454,620)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | (53,333) |  |  |  | (53,333)
Other comprehensive income (loss) | 559 |  |  | 559 |
Exercise of stock options, net (in shares) |  | 702 |  |  |
Exercise of stock options, net | 3,502 | $ 7 | 3,495 |  |
Vesting of restricted stock units (in shares) |  | 99 |  |  |
Vesting of restricted stock units | $ 0 | $ 1 | (1) |  |
Employee stock purchase plan sales, net (in shares) | 176 | 176 |  |  |
Employee stock purchase plan sales | $ 1,575 | $ 2 | 1,573 |  |
Stock-based compensation expense | 14,007 |  | 14,007 |  |
Common stock, shares outstanding, ending balance (in shares) at Mar. 31, 2023 |  | 188,883 |  |  |
Ending balance at Mar. 31, 2023 | $ (328,287) | $ 1,889 | 1,177,192 | 585 | (1,507,953)
Common stock, shares outstanding, beginning balance (in shares) at Dec. 31, 2023 | 205,771 | 205,771 |  |  |
Beginning balance at Dec. 31, 2023 | $ (455,528) | $ 2,058 | 1,222,236 | 1,337 | (1,681,159)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |
Net loss | (35,379) |  |  |  | (35,379)
Other comprehensive income (loss) | (566) |  |  | (566) |
Exercise of stock options, net (in shares) |  | 176 |  |  |
Exercise of stock options, net | 552 | $ 2 | 550 |  |
Vesting of restricted stock units (in shares) |  | 155 |  |  |
Vesting of restricted stock units | 0 | $ 1 | (1) |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) |  | (8) |  |  |
Shares withheld for taxes for vesting of restricted stock units | $ (38) |  | (38) |  |
Employee stock purchase plan sales, net (in shares) | 251 | 251 |  |  |
Employee stock purchase plan sales | $ 1,129 | $ 2 | 1,127 |  |
Issuance of shares to directors in lieu of cash retainer (in shares) |  | 2 |  |  |
Issuance of shares to directors in lieu of cash retainer | 11 |  | 11 |  |
Stock-based compensation expense | $ 13,652 |  | 13,652 |  |
Common stock, shares outstanding, ending balance (in shares) at Mar. 31, 2024 | 206,347 | 206,347 |  |  |
Ending balance at Mar. 31, 2024 | $ (476,167) | $ 2,063 | $ 1,237,537 | $ 771 | $ (1,716,538)

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