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Assets - Fair Value (Details) - USD ($) $ in Thousands | 3 Months Ended | 6 Months Ended |  |  |
--- | --- | --- | --- | --- | ---
Jun. 30, 2017 | Jun. 30, 2017 | Dec. 31, 2016 | Jun. 30, 2016 | Dec. 31, 2015
Cash equivalents |  |  |  |  |
Total | $ 205,564 | $ 205,564 | $ 277,444 | $ 257,641 | $ 269,232
Short-term investments |  |  |  |  |
Total | 97,086 | 97,086 | 66,787 |  |
Transfer of assets between levels | 0 | 0 |  |  |
Assets measured on a recurring basis |  |  |  |  |
Cash equivalents |  |  |  |  |
Total |  |  | 1,501 |  |
Short-term investments |  |  |  |  |
Total | 97,086 | 97,086 | 66,787 |  |
Assets measured on a recurring basis | U.S. treasuries |  |  |  |  |
Short-term investments |  |  |  |  |
Total | 34,951 | 34,951 | 40,008 |  |
Assets measured on a recurring basis | Government agency securities |  |  |  |  |
Short-term investments |  |  |  |  |
Total | 62,135 | 62,135 | 10,012 |  |
Assets measured on a recurring basis | Corporate debt |  |  |  |  |
Cash equivalents |  |  |  |  |
Total |  |  | 1,501 |  |
Short-term investments |  |  |  |  |
Total |  |  | 13,773 |  |
Assets measured on a recurring basis | Commercial paper |  |  |  |  |
Short-term investments |  |  |  |  |
Total |  |  | 2,994 |  |
Assets measured on a recurring basis | Level 1 |  |  |  |  |
Short-term investments |  |  |  |  |
Total | 34,951 | 34,951 | 40,008 |  |
Assets measured on a recurring basis | Level 1 | U.S. treasuries |  |  |  |  |
Short-term investments |  |  |  |  |
Total | 34,951 | 34,951 | 40,008 |  |
Assets measured on a recurring basis | Level 2 |  |  |  |  |
Cash equivalents |  |  |  |  |
Total |  |  | 1,501 |  |
Short-term investments |  |  |  |  |
Total | 62,135 | 62,135 | 26,779 |  |
Assets measured on a recurring basis | Level 2 | Government agency securities |  |  |  |  |
Short-term investments |  |  |  |  |
Total | $ 62,135 | $ 62,135 | 10,012 |  |
Assets measured on a recurring basis | Level 2 | Corporate debt |  |  |  |  |
Cash equivalents |  |  |  |  |
Total |  |  | 1,501 |  |
Short-term investments |  |  |  |  |
Total |  |  | 13,773 |  |
Assets measured on a recurring basis | Level 2 | Commercial paper |  |  |  |  |
Short-term investments |  |  |  |  |
Total |  |  | $ 2,994 |  |

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