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Consolidated Statements of Convertible Preferred Stock and Stockholders' Equity (Deficit) - USD ($) $ in Thousands | Total | Purchase Agreement Gilead | Cumulative Effect, Period of Adoption, Adjustment | Convertible Preferred Stock | Common Stock | Common Stock Purchase Agreement Gilead | Additional Paid-In Capital | Additional Paid-In Capital Purchase Agreement Gilead | Accumulated Deficit | Accumulated Deficit Cumulative Effect, Period of Adoption, Adjustment | Accumulated Other Comprehensive Income (Loss)
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2017 | $ (72,328) |  |  |  |  |  | $ 948 |  | $ (73,234) |  | $ (42)
Balance, shares at Dec. 31, 2017 |  |  |  | 30,459,574 |  |  |  |  |  |  |
Balance at Dec. 31, 2017 |  |  |  | $ 226,196 |  |  |  |  |  |  |
Balance, shares at Dec. 31, 2017 |  |  |  |  | 3,278,129 |  |  |  |  |  |
Conversion of preferred stock to common stock | 226,198 |  |  |  | $ 3 |  | 226,195 |  |  |  |
Conversion of preferred stock to common stock, shares |  |  |  | (30,459,574) |  |  |  |  |  |  |
Conversion of preferred stock to common stock |  |  |  | $ (226,196) |  |  |  |  |  |  |
Conversion of preferred stock to common stock, shares |  |  |  |  | 30,459,574 |  |  |  |  |  |
Issuance of common stock | 124,735 |  |  |  | $ 1 |  | 124,734 |  |  |  |
Issuance of common stock, shares |  |  |  |  | 9,200,000 |  |  |  |  |  |
Issuance of common stock upon exercise of stock options | 95 |  |  |  |  |  | 95 |  |  |  |
Issuance of common stock upon exercise of stock options, shares |  |  |  |  | 67,349 |  |  |  |  |  |
Vesting of early exercised stock options and restricted stock | 1,276 |  |  |  |  |  | 1,276 |  |  |  |
Vesting of early exercised stock options and restricted stock, shares |  |  |  |  | 528,374 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 751 |  |  |  |  |  | 751 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan, shares |  |  |  |  | 77,397 |  |  |  |  |  |
Stock-based compensation | 3,874 |  |  |  |  |  | 3,874 |  |  |  |
Other comprehensive income (loss) | (65) |  |  |  |  |  |  |  |  |  | (65)
Net loss | (49,594) |  |  |  |  |  |  |  | (49,594) |  |
Balance at Dec. 31, 2018 | 234,942 |  |  |  | $ 4 |  | 357,873 |  | (122,828) |  | (107)
Balance, shares at Dec. 31, 2018 |  |  |  |  | 43,610,823 |  |  |  |  |  |
Issuance of common stock upon exercise of stock options | 188 |  |  |  |  |  | 188 |  |  |  |
Issuance of common stock upon exercise of stock options, shares |  |  |  |  | 34,780 |  |  |  |  |  |
Vesting of early exercised stock options and restricted stock | 1,029 |  |  |  |  |  | 1,029 |  |  |  |
Vesting of early exercised stock options and restricted stock, shares |  |  |  |  | 417,883 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 1,029 |  |  |  |  |  | 1,029 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan, shares |  |  |  |  | 148,709 |  |  |  |  |  |
Stock-based compensation | 8,981 |  |  |  |  |  | 8,981 |  |  |  |
Other comprehensive income (loss) | 171 |  |  |  |  |  |  |  |  |  | 171
Net loss | (84,710) |  |  |  |  |  |  |  | (84,710) |  |
Balance at Dec. 31, 2019 | $ 163,842 |  |  |  | $ 4 |  | 369,100 |  | (205,326) |  | 64
Balance (ASC 606) at Dec. 31, 2019 |  |  | $ 2,212 |  |  |  |  |  |  | $ 2,212 |
Balance, shares at Dec. 31, 2019 |  |  |  | 0 |  |  |  |  |  |  |
Balance, shares at Dec. 31, 2019 | 45,925,004 |  |  |  | 44,212,195 |  |  |  |  |  |
Issuance of common stock | $ 326,246 | $ 107,468 |  |  | $ 2 |  | 326,244 | $ 107,468 |  |  |
Issuance of common stock, shares |  |  |  |  | 12,650,000 | 5,963,029 |  |  |  |  |
Issuance of common stock upon exercise of stock options, shares | 392,523 |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock | $ 3,366 |  |  |  |  |  | 3,366 |  |  |  |
Issuance of common stock upon exercise of stock options and vesting of restricted stock, shares |  |  |  |  | 405,752 |  |  |  |  |  |
Vesting of early exercised stock options and restricted stock | 848 |  |  |  |  |  | 848 |  |  |  |
Vesting of early exercised stock options and restricted stock, shares |  |  |  |  | 258,824 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 1,587 |  |  |  |  |  | 1,587 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan, shares |  |  |  |  | 202,101 |  |  |  |  |  |
Stock-based compensation | 21,825 |  |  |  |  |  | 21,825 |  |  |  |
Other comprehensive income (loss) | (20) |  |  |  |  |  |  |  |  |  | (20)
Net loss | (122,858) |  |  |  |  |  |  |  | (122,858) |  |
Balance at Dec. 31, 2020 | $ 502,304 |  |  |  | $ 6 |  | $ 830,438 |  | $ (328,184) |  | $ 44
Balance, shares at Dec. 31, 2020 |  |  |  | 0 |  |  |  |  |  |  |
Balance, shares at Dec. 31, 2020 | 65,114,685 |  |  |  | 63,691,901 |  |  |  |  |  |

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