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License and Collaboration Agreements - Summary of Revenues (Details) - USD ($) $ in Thousands | 3 Months Ended | 12 Months Ended
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Dec. 31, 2020 | Sep. 30, 2020 | Jun. 30, 2020 | Mar. 31, 2020 | Dec. 31, 2019 | Sep. 30, 2019 | Jun. 30, 2019 | Mar. 31, 2019 | Dec. 31, 2020 | Dec. 31, 2019 | Dec. 31, 2018
License And Collaboration Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |
License revenue | $ 9,487 | $ 64,530 | $ 1,750 | $ 1,750 | $ 9,750 | $ 1,750 | $ 1,750 | $ 1,750 |  |  |
Gilead and Taiho | License |  |  |  |  |  |  |  |  |  |  |
License And Collaboration Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |
License revenue |  |  |  |  |  |  |  |  | $ 55,096 | $ 8,000 | $ 3,000
Gilead and Taiho | Collaboration Revenue |  |  |  |  |  |  |  |  |  |  |
License And Collaboration Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |
License revenue |  |  |  |  |  |  |  |  | 22,421 | 7,000 | 5,353
Gilead and Taiho | Collaboration and License |  |  |  |  |  |  |  |  |  |  |
License And Collaboration Agreements [Line Items] |  |  |  |  |  |  |  |  |  |  |
License revenue |  |  |  |  |  |  |  |  | $ 77,517 | $ 15,000 | $ 8,353

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