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Related Parties - Additional Information (Details) - USD ($) $ / shares in Units, $ in Thousands |  | 1 Months Ended | 3 Months Ended |
--- | --- | --- | --- | ---
Jul. 13, 2020 | Jan. 31, 2022 | Feb. 28, 2021 | May 31, 2020 | Mar. 31, 2022 | Mar. 31, 2021 | Dec. 31, 2021
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Receivable from collaboration partners, current |  |  |  |  | $ 29,419 |  | $ 744,595
Related party, deferred revenue - current |  |  |  |  | 86,038 |  | 96,981
Related party, deferred revenue - noncurrent |  |  |  |  | $ 456,905 |  | $ 462,217
Gilead |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Percentage of outstanding common stock held |  |  |  |  | 19.30% |  |
Number of shares issued |  |  |  | 2,200,000 |  |  |
Value of common stock issued |  |  |  | $ 56,700 |  |  |
Option payments received |  | $ 725,000 |  |  |  |  |
Gilead | Operating Expenses |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Reimbursed under cost-sharing provisions of arrangement |  |  |  |  | $ 30,400 | $ 4,900 |
Gilead | Cost Sharing Receivable |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Receivable from collaboration partners, current |  |  |  |  | 29,300 |  |
Related party, deferred revenue - current |  |  |  |  | 86,000 |  |
Related party, deferred revenue - noncurrent |  |  |  |  | 456,900 |  |
Gilead | Cost Sharing Receivable | Other Noncurrent Assets |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Receivable from collaboration partners, noncurrent |  |  |  |  | $ 1,100 |  |
Gilead | Maximum |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Right to purchase additional outstanding voting common stock percentage |  |  |  |  | 35.00% |  |
Gilead | Gilead Collaboration Agreement |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Revenue recognized |  |  |  |  | $ 16,300 | $ 7,700 |
Upfront cash payment |  |  |  |  | 175,000 |  |
Gilead | Stock Purchase Agreement |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |
Issuance of common stock, shares | 5,963,029 |  | 5,650,000 |  |  |  |
Issuance of common stock |  |  | $ 220,400 |  | 327,800 |  |
Funds received for purchase of common stock |  |  |  |  | 200,000 |  |
Fair value of stock purchased |  |  |  |  | 109,400 |  |
Premium on stock purchased |  |  |  |  | $ 90,600 |  |
Shares issued, price per share |  |  | $ 39.00 |  |  |  |

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