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CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2019
Mar. 31, 2018
Operating Activities:    
Net loss $ (43,218) $ (46,352)
Reconciliation of net loss to net cash used in operating activities:    
Depreciation and amortization 1,939 2,079
Loss (Gain) on disposal of property and equipment 88 (47)
Non-cash impact of lease termination 0 (4,381)
Amortization of debt issuance costs 356 356
Non-cash stock-based compensation 5,558 5,245
Other (14) (435)
Changes in operating assets and liabilities:    
Prepaid expenses and other assets (296) 0
Accounts payable and accrued expenses (11,853) (13,293)
Deferred revenue (3,167) (9,247)
Net cash used in operating activities (50,607) (66,075)
Investing Activities:    
Capital expenditures (805) (150)
Proceeds from maturities of marketable securities 22,000 22,721
Purchases of marketable securities (2,484) (1,984)
Net cash provided by investing activities 18,711 20,587
Financing Activities:    
Net proceeds from sales of common stock 55,239 42,599
Proceeds from the exercise of stock options and employee stock purchases 942 1,325
Net cash provided by financing activities 56,181 43,924
Effect of exchange rate on cash, cash equivalents and restricted cash (45) (26)
Net increase (decrease) in cash, cash equivalents and restricted cash 24,240 (1,590)
Cash, cash equivalents and restricted cash at beginning of period 81,959 135,431
Cash, cash equivalents and restricted cash at end of period 106,199 133,841
Supplemental disclosure of non-cash activities:    
Property and equipment purchases included in accounts payable and accrued expenses 194 62
Supplemental disclosure of cash flow information:    
Cash payments of interest $ 6,094 $ 6,094