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Collaboration and Licensing Agreements - Narrative (Details) $ in Millions, ¥ in Billions	1 Months Ended					3 Months Ended	12 Months Ended						50 Months Ended	79 Months Ended
	Nov. 30, 2019 USD ($) item	Mar. 31, 2018 USD ($)	Dec. 31, 2017 USD ($)	Nov. 30, 2017 USD ($)	Jun. 30, 2015 USD ($)	Mar. 31, 2019 USD ($)	Dec. 31, 2021 USD ($)	Dec. 31, 2021 CAD ($)	Dec. 31, 2020 USD ($)	Dec. 31, 2019 USD ($)	Dec. 31, 2018 USD ($)	Dec. 31, 2017 USD ($)	Dec. 31, 2021 USD ($)	Dec. 31, 2021 USD ($)	Dec. 31, 2021 JPY (¥)	Feb. 28, 2019 USD ($)	Jan. 01, 2018 USD ($)
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							$ 10,097,000		$ 7,571,000	$ 5,281,000
Accounts receivable							502,000		0				$ 502,000	$ 502,000
Deferred revenue							0		4,177,000				0	0
Cost of revenue							1,000,000		145,000	600,000
AstraZeneca
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Cost of revenue							1,000,000		145,000	600,000	$ 466,000	$ 9,400,000
Maximum potential payment per agreement							75,000,000						75,000,000	75,000,000
Licensing revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							5,013,000		706,000	4,500,000
Product supply revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							907,000		1,501,000	322,000
2019 KKC Agreement
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							4,200,000		5,400,000
Deferred revenue							0		4,177,000	$ 4,541,000			0	0
2019 KKC Agreement | Kyowa Kirin Co. Ltd
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront license fees	$ 10,000,000
First installment of upfront license fees	$ 5,000,000
Term of payment of license fee, first installment	30 days
Second installment of upfront license fees	$ 5,000,000
Term of agreement	2 years
Initial transaction price	$ 10,000,000
2019 KKC Agreement | Kyowa Kirin Co. Ltd | Minimum
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Number of separate collaborative agreements | item	1
2017 KKC Agreement
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Accounts receivable							500,000						500,000	500,000
Prepayment received							3,200,000
Unbilled prepayments							1,500,000						1,500,000	1,500,000
2017 KKC Agreement | Kyowa Kirin Co. Ltd
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront license fees				$ 30,000,000
Unbilled revenue																$ 0
Potential development milestones				$ 55,000,000
Potential commercialization milestones							73,900,000						73,900,000	73,900,000	¥ 8.5
2017 KKC Agreement | AstraZeneca
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Uncharged license fee reclassified to accounts payable						$ 1,000,000
2017 KKC Agreement | Licensing revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							5,000,000		0				$ 10,000,000
2017 KKC Agreement | Product supply revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							900,000		1,400,000
AstraZeneca Termination Agreement | AstraZeneca
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Cost of revenue							1,000,000		100,000					$ 11,600,000
Percentage of royalty revenue					10.00%
Percentage of non royalty revenue					20.00%
Maximum potential payment per agreement					$ 75,000,000
Fosun Agreement
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							0		0
Fosun Agreement | Fosun Pharma
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront payment received			$ 12,000,000
Potential development and commercialization milestones							$ 110,000,000
Threshold percentage of net sales for tiered royalties							20.00%	20.00%
Knight Agreement | Knight
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront payment received		$ 2,300,000
Potential development and commercialization milestones							$ 17,400,000	$ 22.2
Knight Agreement | Licensing revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							13,000		700,000
Knight Agreement | Product supply revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							0		100,000
XuanZhu Agreement | XuanZhu
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Upfront payment received	$ 800,000
License fee recognized	1,500,000
Second milestone payment	$ 800,000
XuanZhu Agreement | Licensing revenue
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Revenue							$ 0		$ 0
ASU 2014-09 | Adjustments | 2017 KKC Agreement | Kyowa Kirin Co. Ltd
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Unbilled revenue																	$ 5,000,000
ASU 2014-09 | Adjustments | 2017 KKC Agreement | AstraZeneca
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Uncharged license fee																	$ 1,000,000

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