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Related Party Transactions (Details) - USD ($) $ / shares in Units, $ in Thousands | 1 Months Ended | 3 Months Ended | 9 Months Ended |
--- | --- | --- | --- | ---
Sep. 30, 2016 | Jun. 30, 2016 | Nov. 30, 2015 | Sep. 30, 2016 | Sep. 30, 2015 | Sep. 30, 2016 | Sep. 30, 2015 | Dec. 31, 2015
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Shares issued as payment for services |  |  |  |  |  | $ 8,284 |  |
Fair value of financial assets measured at fair value on a recurring basis | $ 376,093 |  |  | $ 376,093 |  | 376,093 |  | $ 291,762
Collaboration and licensing revenues |  |  |  | 30,590 | $ 34,726 | 82,144 | $ 66,690 |
Third Security, LLC |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Expense for services |  |  |  | $ 156 | 142 | $ 301 | 294 |
Initial term of services agreement |  |  | 1 year |  |  |  |  |
Shares issued as payment for services (in shares) |  |  |  | 89,326 |  | 254,496 |  |
Shares issued as payment for services |  |  |  | $ 2,132 |  | $ 6,542 |  |
Third Security, LLC | Common Stock |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Expense for services |  |  | $ 800 |  |  |  |  |
Fibrocell Science, Inc. |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Payments to acquire convertible notes and warrants | $ 2,604 |  |  |  |  |  |  |
Oragenics, Inc. |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Common stock purchased from collaborative partners, shares |  | 2,261,419 |  |  |  |  |  |
Price per share of common shares (usd per share) |  | $ 0.52 |  |  |  |  |  |
Oragenics, Inc. | Maximum |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Maximum percentage of shares of future securities offerings of collaborative partners to which the entity is entitled to purchase | 30.00% |  |  | 30.00% |  | 30.00% |  |
Related Parties, Aggregated |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Collaboration and licensing revenues |  |  |  | $ 26,688 | 31,740 | $ 70,299 | 59,775 |
Harvest Intrexon Enterprise Fund I, LP | Other nonoperating income |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Management fees revenue |  |  |  | 613 | $ 697 | 1,871 | $ 697 |
Convertible Note and Warrants | Other Assets | Fibrocell Science, Inc. |  |  |  |  |  |  |  |
Related Party Transaction [Line Items] |  |  |  |  |  |  |  |
Fair value of financial assets measured at fair value on a recurring basis | $ 1,990 |  |  | $ 1,990 |  | $ 1,990 |  |

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