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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator (Details) - USD ($) $ in Thousands | Sep. 30, 2016 | Dec. 31, 2015
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Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | $ 322,397 | $ 197,729
Upfront and Milestone Payments |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 309,126 | 181,331
Upfront and Milestone Payments | ZIOPHARM Oncology, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 143,651 | 30,338
Upfront and Milestone Payments | Oragenics, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 8,027 | 8,813
Upfront and Milestone Payments | Fibrocell Science, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 19,631 | 21,445
Upfront and Milestone Payments | Genopaver, LLC |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 2,045 | 2,250
Upfront and Milestone Payments | Intrexon Energy Partners, LLC |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 18,750 | 20,625
Upfront and Milestone Payments | Persea Bio, LLC |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 4,125 | 4,500
Upfront and Milestone Payments | Ares Trading S.A. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 48,776 | 53,567
Upfront and Milestone Payments | Thrive Agrobiotics, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 1,483 | 1,621
Upfront and Milestone Payments | Intrexon Energy Partners II |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 16,333 | 17,833
Upfront and Milestone Payments | Exotech Bio, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 4,722 | 0
Upfront and Milestone Payments | Relieve Genetics, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 4,093 | 0
Upfront and Milestone Payments | Intrexon T1D Partners, LLC |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 9,383 | 0
Upfront and Milestone Payments | AD Skincare, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 4,213 | 0
Upfront and Milestone Payments | Genten Therapeutics, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 4,523 | 0
Upfront and Milestone Payments | CRS Bio, Inc. |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | 2,111 | 0
Upfront and Milestone Payments | Other |  |
Deferred Revenue Arrangement [Line Items] |  |
Deferred revenue | $ 17,260 | $ 20,339

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