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Mergers and Acquisitions - Oxitec - Additional Information (Details) - Oxitec
$ in Thousands
1 Months Ended 3 Months Ended 9 Months Ended
Sep. 30, 2015
USD ($)
shares
Sep. 30, 2015
USD ($)
Sep. 30, 2015
USD ($)
Business Acquisition [Line Items]      
Percentage of outstanding common stock acquired 100.00% 100.00% 100.00%
Consideration transferred, shares issued or issuable | shares 1,359,343    
Cash $ 90,199    
Consideration transferred, cash and working capital adjustments $ 9,449    
Equity interest issued or issuable, number of shares withheld as escrow | shares 480,422    
Cash consideration withheld as escrow $ 1,991    
Consideration withheld as escrow, transfer period after closing 18 months    
Consideration transferred $ 146,394    
Acquisition related costs $ 1,675    
Selling, general and administrative      
Business Acquisition [Line Items]      
Acquisition related costs   $ 1,644 $ 1,644