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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator (Details) - USD ($) $ in Thousands | Dec. 31, 2016 | Dec. 31, 2015
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Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 310,142 | $ 197,729
Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 297,867 | 181,331
ZIOPHARM Oncology, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 138,809 | 30,338
Oragenics, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 7,766 | 8,813
Fibrocell Science, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 19,026 | 21,445
Genopaver, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 1,977 | 2,250
Intrexon Energy Partners, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 18,125 | 20,625
Persea Bio, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 4,000 | 4,500
Ares Trading S.A. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 47,178 | 53,567
Thrive Agrobiotics, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 1,436 | 1,621
Intrexon Energy Partners II, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 15,833 | 17,833
Exotech Bio, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 4,583 | 0
Relieve Genetics, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 3,972 | 0
Intrexon T1D Partners, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 8,653 | 0
AD Skincare, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 4,092 | 0
Genten Therapeutics, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 4,176 | 0
CRS Bio, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | 1,949 | 0
Other | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 16,292 | $ 20,339

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- Definition Amount of deferred revenue as of balance sheet date. Deferred revenue represents collections of cash or other assets related to a revenue producing activity for which revenue has not yet been recognized. Generally, an entity records deferred revenue when it receives consideration from a customer before achieving certain criteria that must be met for revenue to be recognized in conformity with GAAP. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 605 -SubTopic 10 -Section S99 -Paragraph 1 -Subparagraph (SAB Topic 13.A.3) -URI http://asc.fasb.org/extlink&oid=65893310&loc=d3e214044-122780 Reference 2: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 605 -SubTopic 10 -Section S99 -Paragraph 1 -Subparagraph (SAB Topic 13.A.4(a).Q1) -URI http://asc.fasb.org/extlink&oid=65893310&loc=d3e214044-122780 + Details
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