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Collaboration and Licensing Revenue (Tables) | 12 Months Ended
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Dec. 31, 2016
Organization, Consolidation and Presentation of Financial Statements [Abstract] |
Summarized Collaboration and Licensing Revenues | The following tables summarize the amounts recorded as revenue in the consolidated statements of operations for each significant collaboration or licensing agreement for the years ended December 31, 2016, 2015 and 2014.
| Year Ended December 31, 2016
| Revenue Recognized From |  | Total
| Upfront and Milestone Payments |  | Research and Development Services |
ZIOPHARM Oncology, Inc. | $ | 11,529 |  |  | $ | 22,307 |  |  | $ | 33,836 |
Oragenics, Inc. | 1,047 |  |  | 1,705 |  |  | 2,752 |
Fibrocell Science, Inc. | 2,419 |  |  | 3,523 |  |  | 5,942 |
Genopaver, LLC | 273 |  |  | 5,844 |  |  | 6,117 |
S & I Ophthalmic, LLC | — |  |  | 6,141 |  |  | 6,141 |
OvaXon, LLC | — |  |  | 2,934 |  |  | 2,934 |
Intrexon Energy Partners, LLC | 2,500 |  |  | 15,052 |  |  | 17,552 |
Persea Bio, LLC | 500 |  |  | 778 |  |  | 1,278 |
Ares Trading S.A. | 6,389 |  |  | 3,803 |  |  | 10,192 |
Thrive Agrobiotics, Inc. | 185 |  |  | 1,529 |  |  | 1,714 |
Intrexon Energy Partners II, LLC | 2,000 |  |  | 1,169 |  |  | 3,169 |
Exotech Bio, Inc. | 417 |  |  | 375 |  |  | 792 |
Relieve Genetics, Inc. | 361 |  |  | 878 |  |  | 1,239 |
Intrexon T1D Partners, LLC | 821 |  |  | 1,087 |  |  | 1,908 |
AD Skincare, Inc. | 241 |  |  | 454 |  |  | 695 |
Genten Therapeutics, Inc. | 122 |  |  | 264 |  |  | 386 |
CRS Bio, Inc. | 57 |  |  | 91 |  |  | 148 |
Other | 5,572 |  |  | 7,504 |  |  | 13,076 |
Total | $ | 34,433 |  |  | $ | 75,438 |  |  | $ | 109,871 |
| Year Ended December 31, 2015
| Revenue Recognized From |  | Total
| Upfront and Milestone Payments |  | Research and Development Services |
ZIOPHARM Oncology, Inc. | $ | 2,855 |  |  | $ | 16,451 |  |  | $ | 19,306 |
Oragenics, Inc. | 5,679 |  |  | 856 |  |  | 6,535 |
Fibrocell Science, Inc. | 6,046 |  |  | 6,133 |  |  | 12,179 |
Genopaver, LLC | 273 |  |  | 3,556 |  |  | 3,829 |
S & I Ophthalmic, LLC | — |  |  | 4,115 |  |  | 4,115 |
OvaXon, LLC | — |  |  | 2,540 |  |  | 2,540 |
Intrexon Energy Partners, LLC | 2,500 |  |  | 10,947 |  |  | 13,447 |
Persea Bio, LLC | 500 |  |  | 741 |  |  | 1,241 |
Ares Trading S.A. | 3,933 |  |  | 795 |  |  | 4,728 |
Thrive Agrobiotics, Inc. | 46 |  |  | 220 |  |  | 266 |
Intrexon Energy Partners II, LLC | 167 |  |  | — |  |  | 167 |
Other | 10,514 |  |  | 8,954 |  |  | 19,468 |
Total | $ | 32,513 |  |  | $ | 55,308 |  |  | $ | 87,821 |
| Year Ended December 31, 2014
| Revenue Recognized From |  | Total
| Upfront and Milestone Payments |  | Research and Development Services |
ZIOPHARM Oncology, Inc. | $ | 2,577 |  |  | $ | 12,044 |  |  | $ | 14,621 |
Oragenics, Inc. | 1,045 |  |  | 598 |  |  | 1,643 |
Fibrocell Science, Inc. | 1,794 |  |  | 4,398 |  |  | 6,192 |
Genopaver, LLC | 273 |  |  | 1,510 |  |  | 1,783 |
S & I Ophthalmic, LLC | — |  |  | 2,832 |  |  | 2,832 |
OvaXon, LLC | — |  |  | 2,799 |  |  | 2,799 |
Intrexon Energy Partners, LLC | 1,875 |  |  | 4,227 |  |  | 6,102 |
Other | 2,061 |  |  | 7,179 |  |  | 9,240 |
Total | $ | 9,625 |  |  | $ | 35,587 |  |  | $ | 45,212 |
Summary of Deferred Revenue | Deferred revenue consists of the following:
| December 31,
| 2016 |  | 2015
Upfront and milestone payments | $ | 297,867 |  |  | $ | 181,331 |
Prepaid research and development services | 6,015 |  |  | 10,938 |
Prepaid product and service revenues | 5,554 |  |  | 4,759 |
Other | 706 |  |  | 701 |
Total | $ | 310,142 |  |  | $ | 197,729 |
Current portion of deferred revenue | $ | 53,364 |  |  | $ | 35,366 |
Long-term portion of deferred revenue | 256,778 |  |  | 162,363 |
Total | $ | 310,142 |  |  | $ | 197,729 |
Summary of Deferred Revenue by Collaborator | The following table summarizes the remaining balance of deferred revenue associated with upfront and milestone payments for each significant collaboration and licensing agreement.
| December 31,
| 2016 |  | 2015
ZIOPHARM Oncology, Inc. | $ | 138,809 |  |  | $ | 30,338 |
Oragenics, Inc. | 7,766 |  |  | 8,813 |
Fibrocell Science, Inc. | 19,026 |  |  | 21,445 |
Genopaver, LLC | 1,977 |  |  | 2,250 |
Intrexon Energy Partners, LLC | 18,125 |  |  | 20,625 |
Persea Bio, LLC | 4,000 |  |  | 4,500 |
Ares Trading S.A. | 47,178 |  |  | 53,567 |
Thrive Agrobiotics, Inc. | 1,436 |  |  | 1,621 |
Intrexon Energy Partners II, LLC | 15,833 |  |  | 17,833 |
Exotech Bio, Inc. | 4,583 |  |  | — |
Relieve Genetics, Inc. | 3,972 |  |  | — |
Intrexon T1D Partners, LLC | 8,653 |  |  | — |
AD Skincare, Inc. | 4,092 |  |  | — |
Genten Therapeutics, Inc. | 4,176 |  |  | — |
CRS Bio, Inc. | 1,949 |  |  | — |
Other | 16,292 |  |  | 20,339 |
Total | $ | 297,867 |  |  | $ | 181,331 |

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