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Goodwill and Intangible Assets, Net | 12 Months Ended
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Dec. 31, 2016
Goodwill and Intangible Assets Disclosure [Abstract] |
Goodwill and Intangible Assets, Net | Goodwill and Intangible Assets, Net The changes in the carrying amount of goodwill for the years ended December 31, 2016 and 2015, are as follows:
| 2016 |  | 2015
Beginning balance | $ | 165,169 |  |  | $ | 101,059 |
Acquisitions | — |  |  | 67,403 |
Foreign currency translation adjustments | (7,994 | ) |  | (3,293 | )
Ending balance | $ | 157,175 |  |  | $ | 165,169 |
No goodwill or accumulated impairment losses existed as of December 31, 2016 and 2015. Intangible assets consist of the following as of December 31, 2016:
| Weighted Average Useful Life (Years) |  | Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | 15.9 |  | $ | 236,401 |  |  | $ | (29,748 | ) |  | $ | 206,653 |
Customer relationships | 6.5 |  | 10,700 |  |  | (4,672 | ) |  | 6,028 |
Trademarks | 9.3 |  | 6,800 |  |  | (1,792 | ) |  | 5,008 |
Covenant not to compete | 2.0 |  | 370 |  |  | (339 | ) |  | 31 |
In-process research and development |  |  | 7,895 |  |  | — |  |  | 7,895 |
Total |  |  | $ | 262,166 |  |  | $ | (36,551 | ) |  | $ | 225,615 |
Intangible assets consist of the following as of December 31, 2015:
| Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | $ | 157,411 |  |  | $ | (17,775 | ) |  | $ | 139,636 |
Customer relationships | 10,700 |  |  | (2,739 | ) |  | 7,961 |
Trademarks | 6,800 |  |  | (1,018 | ) |  | 5,782 |
Covenant not to compete | 384 |  |  | (160 | ) |  | 224 |
In-process research and development | 93,932 |  |  | — |  |  | 93,932 |
Total | $ | 269,227 |  |  | $ | (21,692 | ) |  | $ | 247,535 |
The balance of in-process research and development as of December 31, 2016 includes certain in-process research and development technology acquired in the Company's Oxitec acquisition and amortization will begin once certain regulatory approvals have been obtained for the in-process programs. See Note 3 for additional discussion of in-process research and development technologies placed in service during 2016. Amortization expense was $15,185, $9,871 and $4,237 for the years ended December 31, 2016, 2015 and 2014, respectively. Total amortization expense is estimated to be $18,002 for 2017, $17,391 for 2018, $17,068 for 2019, $16,648 for 2020, $16,431 for 2021, and $132,180 for the cumulative period thereafter.

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- Definition The entire disclosure for the aggregate amount of goodwill and a description of intangible assets, which may include (a) for amortizable intangible assets (also referred to as finite-lived intangible assets), the carrying amount, the amount of any significant residual value, and the weighted-average amortization period, (b) for intangible assets not subject to amortization (also referred to as indefinite-lived intangible assets), the carrying amount, and (c) the amount of research and development assets acquired and written off in the period, including the line item in the income statement in which the amounts written off are aggregated, if not readily apparent from the income statement. Also discloses (a) for amortizable intangibles assets in total and by major class, the gross carrying amount and accumulated amortization, the total amortization expense for the period, and the estimated aggregate amortization expense for each of the five succeeding fiscal years, (b) for intangible assets not subject to amortization the carrying amount in total and by major class, and (c) for goodwill, in total and for each reportable segment, the changes in the carrying amount of goodwill during the period (including the aggregate amount of goodwill acquired, the aggregate amount of impairment losses recognized, and the amount of goodwill included in the gain (loss) on disposal of a reporting unit). If any part of goodwill has not been allocated to a reportable segment, discloses the unallocated amount and the reasons for not allocating. For each impairment loss recognized related to an intangible asset (excluding goodwill), discloses: (a) a description of the impaired intangible asset and the facts and circumstances leading to the impairment, (b) the amount of the impairment loss and the method for determining fair value, (c) the caption in the income statement or the statement of activities in which the impairment loss is aggregated, and (d) the segment in which the impaired intangible asset is reported. For each goodwill impairment loss recognized, discloses: (a) a description of the facts and circumstances leading to the impairment, (b) the amount of the impairment loss and the method of determining the fair value of the associated reporting unit, and (c) if a recognized impairment loss is an estimate not finalized and the reasons why the estimate is not final. May also disclose the nature and amount of any significant adjustments made to a previous estimate of an impairment loss. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -SubTopic 30 -Section 50 -Paragraph 3 -URI http://asc.fasb.org/extlink&oid=66006027&loc=d3e16373-109275 Reference 2: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -SubTopic 30 -Section 50 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=66006027&loc=d3e16265-109275 Reference 3: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -SubTopic 30 -Section 50 -Paragraph 2 -URI http://asc.fasb.org/extlink&oid=66006027&loc=d3e16323-109275 Reference 4: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -SubTopic 20 -Section 50 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=35741047&loc=d3e13816-109267 Reference 5: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -SubTopic 20 -Section 50 -Paragraph 2 -URI http://asc.fasb.org/extlink&oid=35741047&loc=d3e13854-109267 + Details
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