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Mergers and Acquisitions - Trans Ova - Fair Value of Consideration Transferred (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Aug. 31, 2014 | Dec. 31, 2016 | Dec. 31, 2015 | Dec. 31, 2014
Business Acquisition [Line Items] |  |  |  |
Deferred cash consideration |  | $ 0 | $ 1,992 | $ 20,115
Trans Ova Genetics, LC |  |  |  |
Business Acquisition [Line Items] |  |  |  |
Cash | $ 63,625 |  |  |
Common shares | 32,802 |  |  |
Deferred cash consideration | 20,115 |  |  |
Total consideration transferred | 116,542 |  |  |
Fair value of noncontrolling interest | 11,333 |  |  |
Total | $ 127,875 |  |  |

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