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Mergers and Acquisitions - Trans Ova - Additional Information (Details) - USD ($) | 1 Months Ended | 3 Months Ended | 12 Months Ended
--- | --- | --- | ---
Feb. 28, 2015 | Aug. 31, 2014 | Dec. 31, 2016 | Sep. 30, 2016 | Jun. 30, 2016 | Mar. 31, 2016 | Dec. 31, 2015 | Sep. 30, 2015 | Jun. 30, 2015 | Mar. 31, 2015 | Dec. 31, 2016 | Dec. 31, 2015 | Dec. 31, 2014
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred cash consideration |  |  |  |  |  |  |  |  |  |  | $ 0 | $ 1,992,000 | $ 20,115,000
Revenues |  |  | $ 46,002,000 | $ 48,985,000 | $ 52,501,000 | $ 43,438,000 | $ 41,498,000 | $ 53,367,000 | $ 44,891,000 | $ 33,849,000 | 190,926,000 | 173,605,000 | 71,930,000
Net income (loss) |  |  | (44,912,000) | $ (30,011,000) | $ (50,031,000) | $ (65,320,000) | $ (33,275,000) | $ (39,029,000) | $ (41,494,000) | $ 25,804,000 | (190,274,000) | (87,994,000) | (85,616,000)
Trans Ova Genetics, LC |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Percentage of outstanding common stock acquired |  | 100.00% |  |  |  |  |  |  |  |  |  |  |
Business combination, consideration paid, shares issued |  | 1,444,388 |  |  |  |  |  |  |  |  |  |  |
Cash |  | $ 63,625,000 |  |  |  |  |  |  |  |  |  |  |
Deferred cash consideration |  | $ 20,115,000 |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | 78,143,000 | 83,912,000 | 26,352,000
Net income (loss) |  |  |  |  |  |  |  |  |  |  | (9,679,000) | $ (1,531,000) | 2,000
Future contingent payments based on revenue targets |  |  | 3,000,000 |  |  |  |  |  |  |  | 3,000,000 |  |
Current contingent consideration liability |  |  | $ 0 |  |  |  |  |  |  |  | $ 0 |  |
Exemplar Genetics, LLC |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Cash paid to acquire minority interest | $ 1,566,000 |  |  |  |  |  |  |  |  |  |  |  |
Shares issued to acquire minority interest | 307,074 |  |  |  |  |  |  |  |  |  |  |  |
Minimum |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Expected useful life of intangible asset |  |  |  |  |  |  |  |  |  |  | 2 years |  |
Minimum | Trans Ova Genetics, LC |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Expected useful life of intangible asset |  | 3 years |  |  |  |  |  |  |  |  |  |  |
Maximum |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Expected useful life of intangible asset |  |  |  |  |  |  |  |  |  |  | 21 years |  |
Maximum | Trans Ova Genetics, LC |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Expected useful life of intangible asset |  | 9 years |  |  |  |  |  |  |  |  |  |  |
Selling, general and administrative | Trans Ova Genetics, LC |  |  |  |  |  |  |  |  |  |  |  |  |
Business Acquisition [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Business combination, acquisition related cost |  |  |  |  |  |  |  |  |  |  |  |  | $ 713,000

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- Definition Percentage of voting equity interests acquired at the acquisition date in the business combination. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 805 -SubTopic 10 -Section 50 -Paragraph 2 -Subparagraph (c) -URI http://asc.fasb.org/extlink&oid=65885119&loc=d3e1392-128463 + Details
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- Definition This element represents acquisition-related costs incurred to effect a business combination which costs have been expensed during the period. Such costs include finder's fees; advisory, legal, accounting, valuation, and other professional or consulting fees; general administrative costs, including the costs of maintaining an internal acquisitions department; and may include costs of registering and issuing debt and equity securities. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 805 -SubTopic 10 -Section 25 -Paragraph 23 -URI http://asc.fasb.org/extlink&oid=73531572&loc=d3e1043-128460 + Details
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- Definition The cash outflow associated with the acquisition of business during the period. The cash portion only of the acquisition price. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Glossary Investing Activities -URI http://asc.fasb.org/extlink&oid=6516133 Reference 2: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 230 -SubTopic 10 -Section 45 -Paragraph 13 -Subparagraph (b) -URI http://asc.fasb.org/extlink&oid=66902209&loc=d3e3213-108585 + Details
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