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Collaboration and Licensing Revenue - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands | Mar. 31, 2019 | Dec. 31, 2018
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Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 71,191 | $ 69,764
Current portion of deferred revenue | 17,149 | 15,554
Long-term portion of deferred revenue | 54,042 | 54,210
Collaboration and licensing agreements |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | 65,135 | 63,284
Prepaid product and service revenues |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | 3,157 | 2,933
Other |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 2,899 | $ 3,547

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