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Goodwill and Intangible Assets, Net | 3 Months Ended
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Mar. 31, 2019
Goodwill and Intangible Assets Disclosure [Abstract] |
Goodwill and Intangible Assets, Net | Goodwill and Intangible Assets, Net The changes in the carrying amount of goodwill for the three months ended March 31, 2019 are as follows:
Balance at December 31, 2018 | $ | 149,585 |
Foreign currency translation adjustments | 1,170 |
Balance at March 31, 2019 | $ | 150,755 |
The Company had $13,823 of accumulated impairment losses as of March 31, 2019 and December 31, 2018. Intangible assets consist of the following as of March 31, 2019:
| Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | $ | 151,887 |  |  | $ | (37,671 | ) |  | $ | 114,216 |
Customer relationships | 10,700 |  |  | (7,783 | ) |  | 2,917 |
Trademarks | 6,800 |  |  | (3,535 | ) |  | 3,265 |
In-process research and development | 5,470 |  |  | — |  |  | 5,470 |
Total | $ | 174,857 |  |  | $ | (48,989 | ) |  | $ | 125,868 |
Intangible assets consist of the following as of December 31, 2018:
| Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | $ | 152,482 |  |  | $ | (35,133 | ) |  | $ | 117,349 |
Customer relationships | 10,700 |  |  | (7,565 | ) |  | 3,135 |
Trademarks | 6,800 |  |  | (3,341 | ) |  | 3,459 |
In-process research and development | 5,348 |  |  | — |  |  | 5,348 |
Total | $ | 175,330 |  |  | $ | (46,039 | ) |  | $ | 129,291 |
The balance of in-process research and development includes certain in-process research and development technology acquired in the Company's acquisition of Oxitec in September 2015, and amortization will begin once certain regulatory approvals have been obtained for the in-process programs. Amortization expense was $3,004 and $4,926 for the three months ended March 31, 2019 and 2018, respectively.

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- Definition The entire disclosure for goodwill and intangible assets. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -URI http://asc.fasb.org/topic&trid=2144416 + Details
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