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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenues (Details) - USD ($) $ in Thousands | 3 Months Ended
--- | ---
Mar. 31, 2019 | Mar. 31, 2018
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | $ 23,335 | $ 39,666
Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 5,970 | 19,848
ZIOPHARM Oncology, Inc. | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 1,166 | 5,377
Ares Trading S.A. | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 0 | 2,423
Oragenics, Inc. | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 203 | 125
Intrexon T1D Partners, LLC | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 0 | 1,328
Intrexon Energy Partners, LLC | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 977 | 1,197
Intrexon Energy Partners II, LLC | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 504 | 378
Genopaver, LLC | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 294 | 1,315
Fibrocell Science, Inc. | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 383 | 293
Persea Bio, LLC | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | (1,272) | 209
Harvest start-up entities | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | 2,723 | 3,197
Other | Collaboration and licensing agreements |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenues | $ 992 | $ 4,006

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