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Property, Plant and Equipment, Net
3 Months Ended
Mar. 31, 2019
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 
March 31,
2019
 
December 31,
2018
Land and land improvements
$
12,475

 
$
12,490

Buildings and building improvements
20,387

 
20,371

Furniture and fixtures
1,891

 
1,891

Equipment
76,716

 
74,555

Leasehold improvements
31,271

 
28,289

Breeding stock
4,603

 
4,582

Computer hardware and software
11,852

 
11,697

Trees
12,076

 
11,910

Construction and other assets in progress
24,123

 
18,880

 
195,394

 
184,665

Less: Accumulated depreciation and amortization
(59,037
)
 
(55,791
)
Property, plant and equipment, net
$
136,357

 
$
128,874


Depreciation expense was $3,573 and $3,456 for the three months ended March 31, 2019 and 2018, respectively.