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Mergers and Acquisitions - Asset Acquisition of Certain Harvest Entities - Additional Information (Details) - USD ($)	1 Months Ended
	Sep. 30, 2018	Mar. 31, 2019	Dec. 31, 2018	Aug. 31, 2018
Business Acquisition [Line Items]
Investments in affiliates		$ 17,627,000	$ 18,859,000
Deferred revenue		$ 71,191,000	$ 69,764,000
Harvest Intrexon Enterprise Fund I, LP
Business Acquisition [Line Items]
Cash received in asset acquisition	$ 15,500,000
Harvest Intrexon Enterprise Fund I, LP | ActoBio Therapeutics Inc.
Business Acquisition [Line Items]
Long-term debt issued to a related party in an asset acquisition	30,000,000
CRS Bio, Inc.; Genten Therapeutics, Inc.; and Relive Genetics, Inc.
Business Acquisition [Line Items]
Investments in affiliates				$ 4,303,000
Ownership interest		100.00%
Deferred revenue	10,078,000
Write-off of in-process research and development acquired in asset acquisition	$ 8,721,000

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