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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenues (Details) - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2019	Mar. 31, 2018
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	$ 23,335	$ 39,666
Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	5,970	19,848
ZIOPHARM Oncology, Inc. | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	1,166	5,377
Ares Trading S.A. | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	0	2,423
Oragenics, Inc. | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	203	125
Intrexon T1D Partners, LLC | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	0	1,328
Intrexon Energy Partners, LLC | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	977	1,197
Intrexon Energy Partners II, LLC | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	504	378
Genopaver, LLC | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	294	1,315
Fibrocell Science, Inc. | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	383	293
Persea Bio, LLC | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	(1,272)	209
Harvest start-up entities | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	2,723	3,197
Other | Collaboration and licensing agreements
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenues	$ 992	$ 4,006

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