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Lines of Credit and Long-Term Debt - Long-Term Debt - Additional Information (Details)	1 Months Ended			3 Months Ended
	Dec. 31, 2018 USD ($)	Sep. 30, 2018 USD ($)	Jul. 31, 2018 USD ($) day $ / shares	Mar. 31, 2019 USD ($)	Mar. 31, 2018 USD ($)
Debt Instrument [Line Items]
Proceeds from long-term debt, net of issuance costs				$ 376,000	$ 0
Deferred income taxes				508,000	4,074,000
Interest expense				4,311,000	$ 99,000
Long-term debt	$ 211,794,000			214,574,000
Convertible debt
Debt Instrument [Line Items]
Long-term debt	203,391,000			205,829,000
Royalty-based financing
Debt Instrument [Line Items]
Long-term debt	2,085,000			2,151,000
ActoBio Therapeutics Inc. | Convertible debt | Harvest Intrexon Enterprise Fund I, LP
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage		3.00%
Interest expense				225,000
Long-term debt				$ 30,515,000
Trans Ova Genetics, LC | Notes payable to banks | American State Bank
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage				3.95%
Long-term debt				$ 4,382,000
Debt instrument, periodic payment				39,000
AquaBounty Technologies, Inc. | Royalty-based financing | Atlantic Canada Opportunities Agency
Debt Instrument [Line Items]
Long-term debt				$ 2,151,000
Royalty on products, percentage				10.00%
Harvest Intrexon Enterprise Fund I, LP | ActoBio Therapeutics Inc.
Debt Instrument [Line Items]
Long-term debt issued to a related party in an asset acquisition		$ 30,000,000
3.5% Convertible Notes Due 2023
Debt Instrument [Line Items]
Aggregate principal amount			$ 200,000,000	$ 200,000,000
Debt instrument, interest rate, stated percentage			3.50%	3.50%
Proceeds from long-term debt, net of issuance costs			$ 193,958,000
Debt issuance costs			$ 6,042,000
Conversion rate			58.6622
Principal amount used in conversion			$ 1,000
Conversion price (in usd per share) | $ / shares			$ 17.05
Debt instrument redemption price, percentage			100.00%
Carrying value of convertible debt	148,101,000		$ 143,723,000	$ 150,314,000
Equity component of convertible debt, net of issuance costs and deferred taxes			50,235,000
Deferred income taxes			$ 13,367,000
Effective interest rate on convertible notes				11.02%
Convertible notes, unamortized discount and issuance costs				$ 49,686,000
Total interest expense				3,963,000
Interest expense payable in cash				1,750,000
Noncash interest expense				2,213,000
3.5% Convertible Notes Due 2023 | Other Accrued Liabilities
Debt Instrument [Line Items]
Accrued interest payable in cash				$ 1,750,000
3.5% Convertible Notes Due 2023 | Minimum
Debt Instrument [Line Items]
Common stock price trading days | day			20
Common stock price consecutive trading days | day			30
Percentage of common share price over conversion price for conversion			130.00%
3.5% Convertible Notes Due 2023 | Maximum
Debt Instrument [Line Items]
Common stock price trading days | day			5
Common stock price consecutive trading days | day			5
Debt instrument redemption price, percentage			98.00%
Merck Convertible Note
Debt Instrument [Line Items]
Aggregate principal amount	$ 25,000,000
Conversion rate upon IPO	0.90
Interest rate applicable to IPO conversion event	0.05

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