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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Prior Year (Details) - Upfront and Milestone Payments $ in Thousands | Dec. 31, 2018 USD ($)
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Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | $ 63,284 |
ZIOPHARM Oncology, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 1,214 |
Oragenics, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 5,810 |
Intrexon Energy Partners, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 10,267 |
Intrexon Energy Partners II, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 14,060 |
Surterra Holdings, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 0 |
Genopaver, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 1,175 |
Fibrocell Science, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 17,519 |
Persea Bio, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 2,697 |
Harvest start-up entities |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 7,644 | [1]
Other |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | $ 2,898 |
[1] | As of September 30, 2019 and December 31, 2018, the balance of deferred revenue for collaborations with Harvest start-up entities includes: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.

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