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Summary of Significant Accounting Policies - Equity Method Investments - Additional Information (Details) - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
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Sep. 30, 2019 | Sep. 30, 2018 | Sep. 30, 2019 | Sep. 30, 2018
AquaBounty Technologies, Inc. |  |  |  |
Schedule of Equity Method Investments [Line Items] |  |  |  |
Equity method investments, fair value disclosure | $ 16,320 |  | $ 16,320 |
Ownership interest | 38.00% |  | 38.00% |
Unrealized appreciation (depreciation) in fair value of equity securities | $ (3,721) |  | $ 2,081 |
Oragenics, Inc. |  |  |  |
Schedule of Equity Method Investments [Line Items] |  |  |  |
Unrealized appreciation (depreciation) in fair value of equity securities |  | $ (387) |  | $ (1,547)

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- Definition The percentage of ownership of common stock or equity participation in the investee accounted for under the equity method of accounting. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 323 -SubTopic 10 -Section 50 -Paragraph 3 -Subparagraph (a)(1) -URI http://asc.fasb.org/extlink&oid=114001798&loc=d3e33918-111571 + Details
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- Definition The net change in the difference between the fair value and the carrying value, or in the comparative fair values, of investments, not including unrealized gains or losses on securities separately or otherwise categorized as trading, available-for-sale, or held-to-maturity, held at each balance sheet date and included in earnings for the period. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 230 -SubTopic 10 -Section 45 -Paragraph 28 -Subparagraph (b) -URI http://asc.fasb.org/extlink&oid=118932676&loc=d3e3602-108585 + Details
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