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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenues (Details) - USD ($)
$ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2019
Sep. 30, 2018
Sep. 30, 2019
Sep. 30, 2018
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues $ 23,043 $ 32,448 $ 82,364 $ 117,389
Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 6,185 14,324 21,252 51,622
ZIOPHARM Oncology, Inc. | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 431 4,826 2,130 13,626
Ares Trading S.A. | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 0 1,576 0 7,525
Oragenics, Inc. | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 231 705 615 867
Intrexon T1D Partners, LLC | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 0 368 0 2,399
Intrexon Energy Partners, LLC | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 823 1,329 2,596 3,345
Intrexon Energy Partners II, LLC | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 293 754 1,217 1,685
Surterra Holdings, Inc. | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 1,022 0 1,182 0
Genopaver, LLC | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 494 689 1,186 3,076
Fibrocell Science, Inc. | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 402 391 3,247 1,015
Persea Bio, LLC | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues 1,083 199 621 714
Harvest start-up entities | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues [1] 100 2,691 4,862 11,792
Other | Collaboration and licensing agreements        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues $ 1,306 $ 796 $ 3,596 $ 5,578
[1]
For the three and nine months ended September 30, 2019 and 2018, revenues recognized from collaborations with Harvest start-up entities include: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc. For the nine months ended September 30, 2018, revenues recognized from collaborations with Harvest start-up entities also include Genten Therapeutics, Inc. and CRS Bio, Inc.