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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Current Year (Details) - USD ($) $ in Thousands		Sep. 30, 2019	Dec. 31, 2018
Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			$ 63,284
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 74,389
ZIOPHARM Oncology, Inc. | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			1,214
ZIOPHARM Oncology, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		0 years
ZIOPHARM Oncology, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 0
Oragenics, Inc. | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			5,810
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years 8 months 12 days
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 5,463
Intrexon Energy Partners, LLC | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			10,267
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years 6 months
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 8,362
Intrexon Energy Partners II, LLC | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			14,060
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		5 years 2 months 12 days
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 12,843
Surterra Holdings, Inc. | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			0
Surterra Holdings, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		8 years 8 months 12 days
Surterra Holdings, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 13,987
Genopaver, LLC | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			1,175
Genopaver, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years 6 months
Genopaver, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 789
Fibrocell Science, Inc. | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			17,519
Fibrocell Science, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		5 years 1 month 6 days
Fibrocell Science, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 18,102
Persea Bio, LLC | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			2,697
Persea Bio, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		5 years 3 months 18 days
Persea Bio, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 3,553
Harvest start-up entities | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	[1]		7,644
Harvest start-up entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)	[1]	5 years 4 months 24 days
Harvest start-up entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	[1]	$ 6,993
Other | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			$ 2,898
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		1 year 8 months 12 days
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2019-10-01 | Upfront and Milestone Payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 4,297
[1] As of September 30, 2019 and December 31, 2018, the balance of deferred revenue for collaborations with Harvest start-up entities includes: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.

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-Publisher FASB
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