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Lines of Credit and Long-Term Debt - Long-Term Debt - Additional Information (Details)	1 Months Ended			3 Months Ended		9 Months Ended
	Dec. 31, 2018 USD ($)	Sep. 30, 2018 USD ($)	Jul. 31, 2018 USD ($) day $ / shares	Sep. 30, 2019 USD ($)	Sep. 30, 2018 USD ($)	Sep. 30, 2019 USD ($)	Sep. 30, 2018 USD ($)
Debt Instrument [Line Items]
Long-term debt divested from deconsolidation				$ 4,030,000		$ 4,030,000
Proceeds from long-term debt, net of issuance costs						376,000	$ 194,000,000
Equity component of convertible debt, net of issuance costs and deferred taxes					$ 36,868,000		36,868,000
Deferred income taxes				467,000	14,246,000	1,448,000	19,335,000
Accrued interest				1,750,000		1,750,000
Long-term debt issued to a related party in an asset acquisition						0	30,000,000
Interest expense				4,471,000	$ 3,999,000	13,140,000	$ 4,240,000
Long-term debt	$ 211,794,000			215,467,000		215,467,000
Convertible debt
Debt Instrument [Line Items]
Long-term debt	203,391,000			211,037,000		211,037,000
Notes payable
Debt Instrument [Line Items]
Long-term debt	4,551,000			4,204,000		4,204,000
ActoBio Therapeutics Inc. | Convertible debt | Harvest Intrexon Enterprise Fund I, LP
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage		3.00%			3.00%		3.00%
Interest expense				232,000	$ 60,000	684,000	$ 60,000
Long-term debt				$ 30,974,000		$ 30,974,000
Trans Ova Genetics, LC | Notes payable | American State Bank
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage				3.95%		3.95%
Long-term debt				$ 4,180,000		$ 4,180,000
Debt instrument, periodic payment						39,000
Harvest Intrexon Enterprise Fund I, LP | ActoBio Therapeutics Inc.
Debt Instrument [Line Items]
Long-term debt issued to a related party in an asset acquisition		$ 30,000,000
3.5% Convertible Notes Due 2023
Debt Instrument [Line Items]
Aggregate principal amount			$ 200,000,000	$ 200,000,000		$ 200,000,000
Proceeds from long-term debt, net of issuance costs			193,958,000
Debt issuance costs			$ 6,042,000
Debt instrument, interest rate, stated percentage			3.50%	3.50%		3.50%
Conversion rate			58.6622
Principal amount used in conversion			$ 1,000
Conversion price (in usd per share) | $ / shares			$ 17.05
Debt instrument redemption price, percentage			100.00%
Carrying value of convertible debt	148,101,000		$ 143,723,000	$ 155,063,000		$ 155,063,000
Equity component of convertible debt, net of issuance costs and deferred taxes			50,235,000
Deferred income taxes			$ 13,367,000		$ 13,367,000
Effective interest rate on convertible notes				11.02%		11.02%
Convertible notes, unamortized discount and issuance costs				$ 44,937,000		$ 44,937,000
3.5% Convertible Notes Due 2023 | Minimum
Debt Instrument [Line Items]
Common stock price trading days | day			20
Common stock price consecutive trading days | day			30
Percentage of common share price over conversion price for conversion			130.00%
3.5% Convertible Notes Due 2023 | Maximum
Debt Instrument [Line Items]
Common stock price trading days | day			5
Common stock price consecutive trading days | day			5
Debt instrument redemption price, percentage			98.00%
Merck Convertible Note
Debt Instrument [Line Items]
Aggregate principal amount	$ 25,000,000
Conversion rate upon IPO	0.90
Interest rate applicable to IPO conversion event	0.05

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