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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Prior Year (Details) - Upfront and milestone payments $ in Thousands | Dec. 31, 2019 USD ($)
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Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | $ 50,593 |
Oragenics, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 2,864 |
Intrexon Energy Partners, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 8,362 |
Intrexon Energy Partners II, LLC |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 12,843 |
Fibrocell Science, Inc. |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 17,697 |
Harvest start-up entities |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | 6,993 | [1]
Other |  |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |  |
Deferred revenue | $ 1,834 |
[1] | As of June 30, 2020 and December 31, 2019, the balance of deferred revenue for collaborations with Harvest start-up entities includes: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.

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