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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenue (Details) - USD ($) $ in Thousands | 3 Months Ended | 6 Months Ended
--- | --- | ---
Jun. 30, 2020 | Jun. 30, 2019 | Jun. 30, 2020 | Jun. 30, 2019
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | $ 30,424 | $ 32,836 | $ 60,262 | $ 55,421
Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 4,315 | 6,450 | 15,036 | 12,421
ZIOPHARM Oncology, Inc. | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 0 | 533 | 100 | 1,699
Oragenics, Inc. | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 32 | 181 | 230 | 382
Intrexon Energy Partners, LLC | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 0 | 796 | 0 | 1,773
Intrexon Energy Partners II, LLC | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 0 | 420 | 0 | 924
Fibrocell Science, Inc. | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | 4,210 | 2,462 | 14,573 | 2,845
Harvest start-up entities | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers | [1] | 0 | 2,039 | 0 | 4,762
Other | Collaborative Arrangement |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |
Revenues from external customers |  | $ 73 | $ 19 | $ 133 | $ 36
[1] | For the three and six months ended June 30, 2019, revenues recognized from collaborations with Harvest start-up entities include: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.

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