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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenue (Details) - USD ($)
$ in Thousands
 3 Months Ended 6 Months Ended
Jun. 30, 2020
Jun. 30, 2019
Jun. 30, 2020
Jun. 30, 2019
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers $ 30,424 $ 32,836 $ 60,262 $ 55,421
Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 4,315 6,450 15,036 12,421
ZIOPHARM Oncology, Inc. | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 0 533 100 1,699
Oragenics, Inc. | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 32 181 230 382
Intrexon Energy Partners, LLC | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 0 796 0 1,773
Intrexon Energy Partners II, LLC | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 0 420 0 924
Fibrocell Science, Inc. | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers 4,210 2,462 14,573 2,845
Harvest start-up entities | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers [1] 0 2,039 0 4,762
Other | Collaborative Arrangement        
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]        
Revenues from external customers $ 73 $ 19 $ 133 $ 36
[1] For the three and six months ended June 30, 2019, revenues recognized from collaborations with Harvest start-up entities include: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.