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Goodwill and Intangible Assets, Net | 12 Months Ended
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Dec. 31, 2020
Goodwill and Intangible Assets Disclosure [Abstract] |
Goodwill and Intangible Assets, Net | Goodwill and Intangible Assets, Net The changes in the carrying amount of goodwill for the years ended December 31, 2020 and 2019, are as follows:
| 2020 |  | 2019
Beginning of year | $ | 54,119 |  |  | $ | 83,992 |
Impairment | — |  |  | (29,820) |
Foreign currency translation adjustments | 244 |  |  | (53) |
End of year | $ | 54,363 |  |  | $ | 54,119 |
The Company had $43,643 of cumulative impairment losses as of December 31, 2020 and 2019. For the year ended December 31, 2019, during the Company's annual goodwill impairment test, the Company determined it was more-likely-than-not that the fair value of the Trans Ova reporting unit was less than its carrying amount. As a result, the Company compared the carrying amount of the Trans Ova reporting unit to the fair value and determined the carrying amount exceeded the fair value resulting in a $29,642 goodwill impairment charge for the excess carrying value. Additionally, the Company recorded $178 of goodwill impairment in conjunction with the closing of two of its reporting units during the third quarter of 2019. See Note 20 for information regarding goodwill by reportable segment. Intangible assets consist of the following as of December 31, 2020:
| Weighted Average Useful Life (Years) |  | Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | 16.0 |  | $ | 96,927 |  |  | $ | (34,412) |  |  | $ | 62,515 |
Customer relationships | 6.4 |  | 10,850 |  |  | (9,340) |  |  | 1,510 |
Trademarks | 8.4 |  | 5,900 |  |  | (4,529) |  |  | 1,371 |
Total |  |  | $ | 113,677 |  |  | $ | (48,281) |  |  | $ | 65,396 |
Intangible assets consist of the following as of December 31, 2019:
| Gross Carrying Amount |  | Accumulated Amortization |  | Net
Patents, developed technologies and know-how | $ | 90,659 |  |  | $ | (26,619) |  |  | $ | 64,040 |
Customer relationships | 10,700 |  |  | (8,440) |  |  | 2,260 |
Trademarks | 5,900 |  |  | (3,854) |  |  | 2,046 |
Total | $ | 107,259 |  |  | $ | (38,913) |  |  | $ | 68,346 |
In the year ended December 31, 2018, the Company recorded a $16,027 loss related to the abandonment of certain developed technologies that the Company ceased using in the fourth quarter of 2018. The Company does not expect to use these technologies as a defensive asset or market them for sale in the future. Because these technologies were used in combination with other technologies, the identifiable cash flows did not result in an impairment loss; however, because the Company made a decision to abandon the assets, it recorded the charge to research and development expense. Amortization expense was $7,593, $7,920 and $11,666 for the years ended December 31, 2020, 2019 and 2018, respectively. Estimated aggregate amortization expense for definite lived intangible assets is expected to be as follows:
2021 | $ | 7,704 |
2022 | 6,908 |
2023 | 5,742 |
2024 | 5,432 |
2025 | 5,419 |
Thereafter | 34,191 |
Total | $ | 65,396 |

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- Definition The entire disclosure for goodwill and intangible assets. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 350 -URI http://asc.fasb.org/topic&trid=2144416 + Details
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