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Consolidated Statements of Shareholders' and Total Equity - USD ($) $ in Thousands | Total | Cumulative effect of adoption of ASC 606 | Total Precigen Shareholders' Equity | Total Precigen Shareholders' Equity Cumulative effect of adoption of ASC 606 | Common Stock | Additional Paid-in Capital | Accumulated Other Comprehensive Income (Loss) | Accumulated Other Comprehensive Income (Loss) Cumulative effect of adoption of ASC 606 | Accumulated Deficit | Accumulated Deficit Cumulative effect of adoption of ASC 606 | Noncontrolling Interests
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2017 | $ 546,545 | $ 26,507 | $ 533,631 | $ 26,507 | $ 0 | $ 1,397,005 | $ (15,554) | $ (104) | $ (847,820) | $ 26,611 | $ 12,914
Beginning balance, shares at Dec. 31, 2017 |  |  |  |  | 122,087,040 |  |  |  |  |  |
Changes in Stockholders' Equity |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense | 36,296 |  | 36,174 |  |  | 36,174 |  |  |  |  | 122
Shares issued upon vesting of restricted stock units and for exercises of stock options and warrants | 2,336 |  | 297 |  |  | 297 |  |  |  |  | 2,039
Shares issued upon vesting of restricted stock units and for exercises of stock options and warrants, shares |  |  |  |  | 70,159 |  |  |  |  |  |
Shares issued as payment for services | 10,695 |  | 10,695 |  |  | 10,695 |  |  |  |  | 0
Shares issued as payment for services, shares |  |  |  |  | 909,980 |  |  |  |  |  |
Shares and warrants issued in public or private offerings, net of issuance costs | 87,990 |  | 82,374 |  |  | 82,374 |  |  |  |  | 5,616
Shares and warrants issued in public or private offerings, net of issuance costs, shares |  |  |  |  | 6,900,000 |  |  |  |  |  |
Equity component of convertible debt, net of issuance costs and deferred taxes | 36,868 |  | 36,868 |  |  | 36,868 |  |  |  |  | 0
Shares issued pursuant to share lending agreement, shares |  |  |  |  | 7,479,431 |  |  |  |  |  |
Shares issued for reacquired in-process research and development | 159,323 |  | 159,323 |  |  | 159,323 |  |  |  |  | 0
Shares issued for reacquired in-process research and development, shares |  |  |  |  | 22,573,856 |  |  |  |  |  |
Adjustments for noncontrolling interests | 0 |  | (724) |  |  | (724) |  |  |  |  | 724
Net loss | (514,706) |  | (509,336) |  |  |  |  |  | (509,336) |  | (5,370)
Release of cumulative translation adjustment to loss from discontinued operations | 0 |  |  |  |  |  |  |  |  |  |
Other comprehensive income (loss) | (13,132) |  | (12,954) |  |  |  | (12,954) |  |  |  | (178)
Ending balance at Dec. 31, 2018 | 378,722 |  | 362,855 |  | $ 0 | 1,722,012 | (28,612) |  | (1,330,545) |  | 15,867
Ending balance, shares at Dec. 31, 2018 |  |  |  |  | 160,020,466 |  |  |  |  |  |
Changes in Stockholders' Equity |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense | 18,950 |  | 18,881 |  |  | 18,881 |  |  |  |  | 69
Shares issued upon vesting of restricted stock units and for exercises of stock options and warrants | 313 |  | 63 |  |  | 63 |  |  |  |  | 250
Shares issued upon vesting of restricted stock units and for exercises of stock options and warrants, shares |  |  |  |  | 1,028,144 |  |  |  |  |  |
Shares issued for accrued compensation | 1,102 |  | 1,102 |  |  | 1,102 |  |  |  |  |
Shares issued for accrued compensation, shares |  |  |  |  | 150,908 |  |  |  |  |  |
Shares issued as payment for services | 10,446 |  | 10,446 |  |  | 10,446 |  |  |  |  |
Shares issued as payment for services, shares |  |  |  |  | 2,075,362 |  |  |  |  |  |
Shares and warrants issued in public or private offerings, net of issuance costs | 6,611 |  |  |  |  |  |  |  |  |  | 6,611
Adjustments for noncontrolling interests | 0 |  | (456) |  |  | (456) |  |  |  |  | 456
Deconsolidation of subsidiary | (21,672) |  |  |  |  |  |  |  |  |  | (21,672)
Net loss | (323,916) |  | (322,324) |  |  |  |  |  | (322,324) |  | (1,592)
Release of cumulative translation adjustment to loss from discontinued operations | 0 |  |  |  |  |  |  |  |  |  |
Other comprehensive income (loss) | 1,155 |  | 1,144 |  |  |  | 1,144 |  |  |  | 11
Ending balance at Dec. 31, 2019 | $ 71,711 |  | 71,711 |  | $ 0 | 1,752,048 | (27,468) |  | (1,652,869) |  | $ 0
Ending balance, shares at Dec. 31, 2019 | 163,274,880 |  |  |  | 163,274,880 |  |  |  |  |  |
Changes in Stockholders' Equity |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  |  | 18,366 |  |  | 18,366 |  |  |  |  |
Shares issued upon vesting of restricted stock units and for exercises of stock options |  |  | 117 |  |  | 117 |  |  |  |  |
Shares issued upon vesting of restricted stock units and for exercises of stock options, shares |  |  |  |  | 877,249 |  |  |  |  |  |
Shares issued for accrued compensation |  |  | 5,100 |  |  | 5,100 |  |  |  |  |
Shares issued for accrued compensation, shares |  |  |  |  | 1,955,405 |  |  |  |  |  |
Shares issued as payment for services |  |  | 1,006 |  |  | 1,006 |  |  |  |  |
Shares issued as payment for services, shares |  |  |  |  | 413,911 |  |  |  |  |  |
Shares and warrants issued in public or private offerings, net of issuance costs |  |  | 35,000 |  |  | 35,000 |  |  |  |  |
Shares and warrants issued in public or private offerings, net of issuance costs, shares |  |  |  |  | 5,972,696 |  |  |  |  |  |
Shares issued upon conversion of long-term debt |  |  | 56,827 |  |  | 56,827 |  |  |  |  |
Shares issued upon conversion of long-term debt, shares |  |  |  |  | 13,051,802 |  |  |  |  |  |
Shares issued in conjunction with settlement agreement |  |  | 18,103 |  |  | 18,103 |  |  |  |  |
Stock issued in conjunction with settlement agreement, shares |  |  |  |  | 2,117,264 |  |  |  |  |  |
Net loss | $ (170,521) |  | (170,521) |  |  |  |  |  | (170,521) |  |
Release of cumulative translation adjustment to loss from discontinued operations | $ 26,957 |  | 26,957 |  |  |  | 26,957 |  |  |  |
Other comprehensive income (loss) |  |  | 4,508 |  |  |  | 4,508 |  |  |  |
Ending balance at Dec. 31, 2020 |  |  | $ 67,174 |  | $ 0 | $ 1,886,567 | $ 3,997 |  | $ (1,823,390) |  |
Ending balance, shares at Dec. 31, 2020 | 187,663,207 |  |  |  | 187,663,207 |  |  |  |  |  |

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