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Mergers and Acquisitions - Asset Acquisition of Certain Harvest Entities (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Sep. 30, 2018 | Dec. 31, 2020 | Dec. 31, 2019 | Dec. 31, 2018
Business Acquisition [Line Items] |  |  |  |
Long-term debt issued to a related party in an asset acquisition |  | $ 0 | $ 0 | $ 30,000
Cash received in asset acquisition |  | 0 | 0 | 15,500
Investments in affiliates |  | 0 | 1,461 |
Deferred revenue |  | 25,823 | 53,833 |
Reacquisition of in-process research and development |  | $ 0 | $ 0 | $ 236,748
Harvest Intrexon Enterprise Fund I, LP |  |  |  |
Business Acquisition [Line Items] |  |  |  |
Cash received in asset acquisition | $ 15,500 |  |  |
Precigen ActoBio, Inc. | Harvest Intrexon Enterprise Fund I, LP |  |  |  |
Business Acquisition [Line Items] |  |  |  |
Long-term debt issued to a related party in an asset acquisition | 30,000 |  |  |
CRS Bio, Inc.; Genten Therapeutics, Inc.; and Relive Genetics, Inc. |  |  |  |
Business Acquisition [Line Items] |  |  |  |
Investments in affiliates | $ 4,303 |  |  |
Ownership interest | 100.00% |  |  |
Deferred revenue | $ 10,078 |  |  |
Reacquisition of in-process research and development | $ 8,721 |  |  |

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