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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Prior Year (Details) - Upfront and Milestone Payments $ in Thousands | Dec. 31, 2019 USD ($)
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Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | $ 50,593
Oragenics, Inc. |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | 2,864
Intrexon Energy Partners, LLC |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | 8,362
Intrexon Energy Partners II, LLC |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | 12,843
Castle Creek Biosciences, Inc. |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | 17,697
Harvest Start-Up Entities |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | 6,993
Other |
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction [Line Items] |
Deferred revenue | $ 1,834

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