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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Current Year (Details) - USD ($) $ in Thousands | Dec. 31, 2020 | Dec. 31, 2019
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Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | $ 50,593
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 23,420 |
Oragenics, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | 2,864
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 0 years |
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 0 |
Intrexon Energy Partners, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | 8,362
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 3 years 2 months 12 days |
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 8,362 |
Intrexon Energy Partners II, LLC | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | 12,843
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 3 years 10 months 24 days |
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 12,843 |
Castle Creek Biosciences, Inc. | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | 17,697
Castle Creek Biosciences, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 1 year |
Castle Creek Biosciences, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 379 |
Harvest Start-Up Entities | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | 6,993
Harvest Start-Up Entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 0 years |
Harvest Start-Up Entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 0 |
Other | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue |  | $ 1,834
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Average Remaining Performance Period (Years) | 2 years 2 months 12 days |
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-01-01 | Upfront and Milestone Payments |  |
Summary of Deferred Revenue by Collaborator [Line Items] |  |
Deferred revenue | $ 1,836 |

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