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Shareholders' Equity - Additional Information (Details) - USD ($) $ / shares in Units, $ in Thousands |  | 1 Months Ended | 12 Months Ended
--- | --- | --- | ---
Jan. 31, 2020 | Mar. 31, 2019 | Oct. 31, 2018 | Jul. 31, 2018 | Jan. 31, 2018 | Dec. 31, 2019 | Dec. 31, 2018
Class of Stock [Line Items] |  |  |  |  |  |  |
Shares issued in public or private offerings, net of issuance costs, shares |  |  |  |  | 6,900,000 |  |
Value of shares issued in public or private offering |  |  |  |  | $ 82,374 | $ 6,611 | $ 87,990
Underwriting discounts and commissions |  |  |  |  | 3,688 |  |
Capitalized offering expenses |  |  |  |  | $ 188 |  |
Shares issued in share lending agreement, shares |  |  |  | 7,479,431 |  |  |
Share lending agreement, length of time for shares to be returned upon termination |  |  |  | 5 days |  |  |
Shares issued, price per share |  |  |  | $ 13.37 |  |  |
Proceeds from shares issued for exercises of warrants |  |  |  |  |  | $ 313 | $ 2,336
TS Biotechnology Sale |  |  |  |  |  |  |
Class of Stock [Line Items] |  |  |  |  |  |  |
Shares issued in public or private offerings, net of issuance costs, shares | 5,972,696 |  |  |  |  |  |
Value of shares issued in public or private offering | $ 35,000 |  |  |  |  |  |
Affiliates of Third Security |  |  |  |  |  |  |
Class of Stock [Line Items] |  |  |  |  |  |  |
Shares issued in public or private offerings, net of issuance costs, shares |  |  |  |  | 1,000,000 |  |
AquaBounty Technologies, Inc. |  |  |  |  |  |  |
Class of Stock [Line Items] |  |  |  |  |  |  |
Value of shares issued in public or private offering |  | $ 6,611 |  |  | $ 10,616 |  |
Proceeds from shares issued for exercises of warrants |  |  | $ 4,316 |  |  |  |
AquaBounty Technologies, Inc. |  |  |  |  |  |  |
Class of Stock [Line Items] |  |  |  |  |  |  |
Additional investment in subsidiary |  |  | $ 3,077 |  | $ 5,000 |  |

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