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Investments in Joint Ventures - Intrexon T1D Partners - Additional Information (Details) | 1 Months Ended | 12 Months Ended
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Nov. 30, 2018 USD ($) shares | Mar. 31, 2016 USD ($) board_seat | Dec. 31, 2020 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($)
Schedule Of Investments In Joint Venture [Line Items] |  |  |  |  |
Shares issued for reacquired in-process research and development |  |  |  |  | $ 159,323,000
Deferred revenue |  |  | $ 25,823,000 | $ 53,833,000 |
Reacquisition of in-process research and development |  |  | 0 | 0 | $ 236,748,000
Collaboration and licensing agreements |  |  |  |  |
Schedule Of Investments In Joint Venture [Line Items] |  |  |  |  |
Deferred revenue |  |  | $ 23,420,000 | $ 50,593,000 |
Intrexon T1D Partners, LLC |  |  |  |  |
Schedule Of Investments In Joint Venture [Line Items] |  |  |  |  |
Membership interest | 100.00% | 50.00% |  |  |
Maximum additional capital contributions committed |  | $ 5,000,000 |  |  |
Total number of seats on the joint venture's governing board, internally selected | board_seat |  | 2 |  |  |
Total number of seats on the joint venture's governing board, externally selected | board_seat |  | 3 |  |  |
Shares issued for reacquired in-process research and development, shares | shares | 1,933,737 |  |  |  |
Shares issued for reacquired in-process research and development | $ 18,970,000 |  |  |  |
Deferred revenue | 8,517,000 |  |  |  |
Reacquisition of in-process research and development | $ 10,453,000 |  |  |  |
Investor | Intrexon T1D Partners, LLC |  |  |  |  |
Schedule Of Investments In Joint Venture [Line Items] |  |  |  |  |
Membership interest |  | 50.00% |  |  |
Capital contribution |  | $ 10,000,000 |  |  |
Maximum additional capital contributions committed |  | 5,000,000 |  |  |
Intrexon T1D Partners, LLC | Collaboration and licensing agreements |  |  |  |  |
Schedule Of Investments In Joint Venture [Line Items] |  |  |  |  |
Collaborative agreement, consideration received, value |  | $ 10,000,000 |  |  |

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