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Segments - Information by Reportable Segment (Details) - USD ($) $ in Thousands	3 Months Ended								12 Months Ended
	Dec. 31, 2020	Sep. 30, 2020	Jun. 30, 2020	Mar. 31, 2020	Dec. 31, 2019	Sep. 30, 2019	Jun. 30, 2019	Mar. 31, 2019	Dec. 31, 2020	Dec. 31, 2019	Dec. 31, 2018
Goodwill
Goodwill	$ 54,363				$ 54,119				$ 54,363	$ 54,119
Impairments						$ (178)			(9,635)	(87,862)	$ 0
Revenues from external customers	19,333	$ 23,583	$ 30,424	$ 29,838	17,002	$ 18,299	$ 32,836	$ 22,585	103,178	90,722	151,178
Segment Adjusted EBITDA									(28,713)	(56,115)	(62,067)
PGEN Therapeutics
Goodwill
Segment Adjusted EBITDA									(25,611)	(30,164)	(32,832)
ActoBio
Goodwill
Segment Adjusted EBITDA									(7,861)	(13,662)	(12,797)
Trans Ova
Goodwill
Impairments										(29,642)
Segment Adjusted EBITDA									2,624	(6,337)	(5,730)
Human Biotherapeutics
Goodwill
Segment Adjusted EBITDA									(2,139)	(1,051)	0
All Other
Goodwill
Segment Adjusted EBITDA									4,274	(4,901)	(10,708)
Operating segments
Goodwill
Goodwill	54,363				54,119				54,363	54,119	83,992
Reallocations from changes to reporting units										0
Impairments										(29,820)
Foreign currency translation adjustments									244	(53)
Revenues from external customers									103,124	86,762	141,096
Revenues									108,748	101,232	143,366
Operating segments | PGEN Therapeutics
Goodwill
Goodwill	15,232				15,232				15,232	15,232	15,232
Reallocations from changes to reporting units										0
Impairments										0
Foreign currency translation adjustments									0	0
Revenues from external customers									776	2,227	29,021
Revenues									5,782	13,568	29,638
Operating segments | ActoBio
Goodwill
Goodwill	1,979				1,735				1,979	1,735	1,788
Reallocations from changes to reporting units										0
Impairments										0
Foreign currency translation adjustments									244	(53)
Revenues from external customers									3,053	(364)	6,684
Revenues									3,050	134	7,524
Operating segments | Trans Ova
Goodwill
Goodwill	16,594				16,594				16,594	16,594	46,236
Reallocations from changes to reporting units										0
Impairments										(29,642)
Foreign currency translation adjustments									0	0
Revenues from external customers									71,186	68,672	75,178
Revenues									71,526	70,033	75,736
Operating segments | Human Biotherapeutics
Goodwill
Goodwill	482				482				482	482	0
Reallocations from changes to reporting units										482
Impairments										0
Foreign currency translation adjustments									0	0
Revenues from external customers									17,810	3,713	0
Revenues									17,810	3,713	0
Operating segments | All Other
Goodwill
Goodwill	$ 20,076				$ 20,076				20,076	20,076	20,736
Reallocations from changes to reporting units										(482)
Impairments										(178)
Foreign currency translation adjustments									0	0
Revenues from external customers									10,299	12,514	30,213
Revenues									10,580	13,784	30,468
Intersegment revenues
Goodwill
Revenues									5,624	14,470	2,270
Intersegment revenues | PGEN Therapeutics
Goodwill
Revenues									5,006	11,341	617
Intersegment revenues | ActoBio
Goodwill
Revenues									(3)	498	840
Intersegment revenues | Trans Ova
Goodwill
Revenues									340	1,361	558
Intersegment revenues | Human Biotherapeutics
Goodwill
Revenues									0	0	0
Intersegment revenues | All Other
Goodwill
Revenues									$ 281	$ 1,270	$ 255

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