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Property, Plant and Equipment, Net
 12 Months Ended
Dec. 31, 2020
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 December 31,
 20202019
Land and land improvements$9,844 $9,814 
Buildings and building improvements12,088 11,765 
Furniture and fixtures1,228 1,025 
Equipment31,150 33,707 
Leasehold improvements6,260 6,607 
Breeding stock868 5,191 
Computer hardware and software5,684 6,877 
Construction and other assets in progress2,754 5,229 
69,876 80,215 
Less: Accumulated depreciation and amortization(34,952)(36,263)
Property, plant and equipment, net$34,924 $43,952 
Depreciation expense was $7,449, $8,222 and $8,946 for the years ended December 31, 2020, 2019 and 2018, respectively.
During the year ended December 31, 2020, the Company recorded impairment losses of $920, which is included in impairment of other noncurrent assets on the accompanying consolidated statement of operations, primarily related to right-of-use assets at certain of the Company's leased locations.
During the year ended December 31, 2019, the Company recorded impairment losses of $448, which is included in impairment of other noncurrent assets on the accompanying consolidated statement of operations, related to the impairment of property, plant and equipment in conjunction with the closing of two of its operating units during the third quarter of 2019.