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Goodwill and Intangible Assets, Net
12 Months Ended
Dec. 31, 2020
Goodwill and Intangible Assets Disclosure [Abstract]  
Goodwill and Intangible Assets, Net Goodwill and Intangible Assets, Net
The changes in the carrying amount of goodwill for the years ended December 31, 2020 and 2019, are as follows:
20202019
Beginning of year$54,119 $83,992 
Impairment— (29,820)
Foreign currency translation adjustments244 (53)
End of year$54,363 $54,119 
The Company had $43,643 of cumulative impairment losses as of December 31, 2020 and 2019.
For the year ended December 31, 2019, during the Company's annual goodwill impairment test, the Company determined it was more-likely-than-not that the fair value of the Trans Ova reporting unit was less than its carrying amount. As a result, the Company compared the carrying amount of the Trans Ova reporting unit to the fair value and determined the carrying amount exceeded the fair value resulting in a $29,642 goodwill impairment charge for the excess carrying value. Additionally, the Company recorded $178 of goodwill impairment in conjunction with the closing of two of its reporting units during the third quarter of 2019.
See Note 20 for information regarding goodwill by reportable segment.
Intangible assets consist of the following as of December 31, 2020: 
Weighted Average Useful Life (Years)Gross Carrying AmountAccumulated AmortizationNet
Patents, developed technologies and know-how16.0$96,927 $(34,412)$62,515 
Customer relationships6.410,850 (9,340)1,510 
Trademarks8.45,900 (4,529)1,371 
Total$113,677 $(48,281)$65,396 
Intangible assets consist of the following as of December 31, 2019: 
Gross Carrying AmountAccumulated AmortizationNet
Patents, developed technologies and know-how$90,659 $(26,619)$64,040 
Customer relationships10,700 (8,440)2,260 
Trademarks5,900 (3,854)2,046 
Total$107,259 $(38,913)$68,346 
In the year ended December 31, 2018, the Company recorded a $16,027 loss related to the abandonment of certain developed technologies that the Company ceased using in the fourth quarter of 2018. The Company does not expect to use these technologies as a defensive asset or market them for sale in the future. Because these technologies were used in combination with other technologies, the identifiable cash flows did not result in an impairment loss; however, because the Company made a decision to abandon the assets, it recorded the charge to research and development expense.
Amortization expense was $7,593, $7,920 and $11,666 for the years ended December 31, 2020, 2019 and 2018, respectively. Estimated aggregate amortization expense for definite lived intangible assets is expected to be as follows:
2021$7,704 
20226,908 
20235,742 
20245,432 
20255,419 
Thereafter34,191 
Total$65,396