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Collaboration and Licensing Revenue (Tables)
12 Months Ended
Dec. 31, 2020
Organization, Consolidation and Presentation of Financial Statements [Abstract]  
Summarized Collaboration and Licensing Revenues
The following table summarizes the amounts recorded as revenue in the consolidated statements of operations for each significant counterparty to a collaboration or licensing agreement for the years ended December 31, 2020, 2019 and 2018.
 Year Ended December 31,
202020192018
ZIOPHARM Oncology, Inc.$200 $2,171 $16,298 
Ares Trading S.A.— — 11,175 
Oragenics, Inc.3,053 (564)1,353 
Intrexon T1D Partners, LLC— — 2,502 
Intrexon Energy Partners, LLC— 2,596 6,929 
Intrexon Energy Partners II, LLC— 1,217 2,998 
Castle Creek Biosciences, Inc.17,810 3,713 1,394 
Harvest start-up entities (1)— 4,862 14,447 
Other145 64 12,444 
Total (2)$21,208 $14,059 $69,540 
(1)For the years ended December 31, 2019 and 2018, revenue recognized from collaborations with Harvest start-up entities include Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc. For the year ended December 31, 2018, revenue recognized from collaborations with Harvest start-up entities also include Genten Therapeutics and CRS Bio.
(2)Collaboration and licensing revenues recognized for the years ended December 31, 2020, 2019 and 2018, include the recognition of $20,205, $7,505, and $39,446, respectively, associated with upfront and milestone payments which were previously deferred.
Summary of Deferred Revenue Deferred revenue consisted of the following:
 December 31,
 20202019
Collaboration and licensing agreements$23,420 $50,593 
Prepaid product and service revenues2,126 2,805 
Other277 435 
Total$25,823 $53,833 
Current portion of deferred revenue$2,800 $5,697 
Long-term portion of deferred revenue23,023 48,136 
Total$25,823 $53,833 
Summary of Deferred Revenue by Collaborator The following table summarizes the remaining balance of deferred revenue associated with upfront and milestone payments for each significant counterparty to a collaboration or licensing agreement as of December 31, 2020 and 2019, as well as the estimated remaining performance period as of December 31, 2020.
Average Remaining Performance Period (Years)December 31,
20202019
Oragenics, Inc.0.0$— $2,864 
Intrexon Energy Partners, LLC3.28,362 8,362 
Intrexon Energy Partners II, LLC3.912,843 12,843 
Castle Creek Biosciences, Inc.1.0379 17,697 
Harvest start-up entities (1)0.0— 6,993 
Other2.21,836 1,834 
Total$23,420 $50,593 
(1)As of December 31, 2019, the balance of deferred revenue for collaborations with Harvest start-up entities includes Thrive Agrobiotics, Exotech Bio, and AD Skincare.