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Mergers and Acquisitions - Asset Acquisition of Certain Harvest Entities (Details) - USD ($)
$ in Thousands
1 Months Ended 12 Months Ended
Sep. 30, 2018
Dec. 31, 2020
Dec. 31, 2019
Dec. 31, 2018
Business Acquisition [Line Items]        
Long-term debt issued to a related party in an asset acquisition   $ 0 $ 0 $ 30,000
Cash received in asset acquisition   0 0 15,500
Investments in affiliates   0 1,461  
Deferred revenue   25,823 53,833  
Reacquisition of in-process research and development   $ 0 $ 0 $ 236,748
Harvest Intrexon Enterprise Fund I, LP        
Business Acquisition [Line Items]        
Cash received in asset acquisition $ 15,500      
Precigen ActoBio, Inc. | Harvest Intrexon Enterprise Fund I, LP        
Business Acquisition [Line Items]        
Long-term debt issued to a related party in an asset acquisition 30,000      
CRS Bio, Inc.; Genten Therapeutics, Inc.; and Relive Genetics, Inc.        
Business Acquisition [Line Items]        
Investments in affiliates $ 4,303      
Ownership interest 100.00%      
Deferred revenue $ 10,078      
Reacquisition of in-process research and development $ 8,721