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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenues (Details) - USD ($)
$ in Thousands
 1 Months Ended 3 Months Ended 12 Months Ended
Jul. 31, 2020
Feb. 29, 2020
Oct. 31, 2018
Dec. 31, 2020
Sep. 30, 2020
Jun. 30, 2020
Mar. 31, 2020
Dec. 31, 2019
Sep. 30, 2019
Jun. 30, 2019
Mar. 31, 2019
Dec. 31, 2020
Dec. 31, 2019
Dec. 31, 2018
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues       $ 19,333 $ 23,583 $ 30,424 $ 29,838 $ 17,002 $ 18,299 $ 32,836 $ 22,585 $ 103,178 $ 90,722 $ 151,178
Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       21,208 14,059 69,540
Upfront and Milestone Payments                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenue recognized from previously deferred balances                       20,205 7,505 39,446
ZIOPHARM Oncology, Inc. | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues     $ 500                 200 2,171 16,298
Ares Trading S.A. | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       0 0 11,175
Oragenics, Inc. | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       3,053 (564) 1,353
Revenue recognized from previously deferred balances $ 2,823                          
Intrexon T1D Partners, LLC | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       0 0 2,502
Intrexon Energy Partners, LLC | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       0 2,596 6,929
Intrexon Energy Partners II, LLC | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       0 1,217 2,998
Castle Creek Biosciences, Inc. | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       17,810 3,713 1,394
Revenue recognized from previously deferred balances   $ 10,000                        
Harvest Start-Up Entities | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       0 4,862 14,447
Other revenues | Collaboration and licensing agreements                            
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]                            
Revenues                       $ 145 $ 64 $ 12,444