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Shareholders' Equity - Additional Information (Details) - USD ($)
$ / shares in Units, $ in Thousands
 1 Months Ended 12 Months Ended
Jan. 31, 2020
Mar. 31, 2019
Oct. 31, 2018
Jul. 31, 2018
Jan. 31, 2018
Dec. 31, 2019
Dec. 31, 2018
Class of Stock [Line Items]              
Shares issued in public or private offerings, net of issuance costs, shares         6,900,000    
Value of shares issued in public or private offering         $ 82,374 $ 6,611 $ 87,990
Underwriting discounts and commissions         3,688    
Capitalized offering expenses         $ 188    
Shares issued in share lending agreement, shares       7,479,431      
Share lending agreement, length of time for shares to be returned upon termination       5 days      
Shares issued, price per share       $ 13.37      
Proceeds from shares issued for exercises of warrants           $ 313 $ 2,336
TS Biotechnology Sale              
Class of Stock [Line Items]              
Shares issued in public or private offerings, net of issuance costs, shares 5,972,696            
Value of shares issued in public or private offering $ 35,000            
Affiliates of Third Security              
Class of Stock [Line Items]              
Shares issued in public or private offerings, net of issuance costs, shares         1,000,000    
AquaBounty Technologies, Inc.              
Class of Stock [Line Items]              
Value of shares issued in public or private offering   $ 6,611     $ 10,616    
Proceeds from shares issued for exercises of warrants     $ 4,316        
AquaBounty Technologies, Inc.              
Class of Stock [Line Items]              
Additional investment in subsidiary     $ 3,077   $ 5,000