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Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator (Details) - USD ($) $ in Thousands | Sep. 30, 2021 | Dec. 31, 2020
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Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue |  | $ 23,420
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 23,040 |
Intrexon Energy Partners, LLC | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue |  | 8,362
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Average Remaining Performance Period (Years) | 2 years 6 months |
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 8,362 |
Intrexon Energy Partners II, LLC | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue |  | 12,843
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Average Remaining Performance Period (Years) | 3 years 2 months 12 days |
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 12,843 |
Castle Creek Biosciences, Inc. | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue |  | 379
Castle Creek Biosciences, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Average Remaining Performance Period (Years) | 3 months 18 days |
Castle Creek Biosciences, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 17 |
Other | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue |  | $ 1,836
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Average Remaining Performance Period (Years) | 3 years 1 month 6 days |
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2021-10-01 | Upfront and milestone payments |  |
Contract With Customer, Asset And Liability [Line Items] |  |
Deferred revenue | $ 1,818 |

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