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Collaboration and Licensing Revenue (Tables)
9 Months Ended
Sep. 30, 2021
Organization, Consolidation and Presentation of Financial Statements [Abstract]  
Summarized Collaboration and Licensing Revenues
The following table summarizes the amounts recorded as revenue in the condensed consolidated statements of operations for each significant counterparty to a collaboration or licensing agreement for the three and nine months ended September 30, 2021 and 2020.
 Three Months Ended 
 September 30,
Nine Months Ended 
 September 30,
2021202020212020
ZIOPHARM Oncology, Inc.$— $— $— $100 
Oragenics, Inc.— 2,823 — 3,053 
Castle Creek Biosciences, Inc.18 2,394 371 16,967 
Other18 139 
Total (1)$22 $5,223 $389 $20,259 
(1)Collaboration and licensing revenues include the recognition of $22 and $5,213 for the three months ended September 30, 2021 and 2020, respectively, and $380 and $19,654 for the nine months ended September 30, 2021 and 2020, respectively, associated with upfront and milestone payments which were previously deferred.
Summary of Deferred Revenue Deferred revenue consisted of the following:
September 30,
2021
December 31,
2020
Collaboration and licensing agreements$23,040 $23,420 
Prepaid product and service revenues3,493 2,126 
Other183 277 
Total$26,716 $25,823 
Current portion of deferred revenue$3,693 $2,800 
Long-term portion of deferred revenue23,023 23,023 
Total$26,716 $25,823 
Summary of Deferred Revenue by Collaborator The following table summarizes the remaining balance of deferred revenue associated with upfront and milestone payments for each significant counterparty to a collaboration or licensing agreement as of September 30, 2021 and December 31, 2020, as well as the estimated remaining performance period as of September 30, 2021.
Average Remaining Performance Period (Years)September 30,
2021
December 31,
2020
Intrexon Energy Partners, LLC2.5$8,362 $8,362 
Intrexon Energy Partners II, LLC3.212,843 12,843 
Castle Creek Biosciences, Inc.0.317 379 
Other3.11,818 1,836 
Total$23,040 $23,420