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Collaboration and Licensing Revenue - Narrative (Details) - USD ($) | 1 Months Ended |  |
--- | --- | --- | ---
Jul. 31, 2022 | Sep. 30, 2022 | Dec. 31, 2021
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |
Deferred revenue, net of current portion, related party |  | $ 1,818,000 | $ 23,023,000
Intrexon Energy Partners L.L. and Intrexon Energy Partners II, L.L.C |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |
Deferred revenue, net of current portion, related party |  | $ 0 | $ 21,205,000
Intrexon Energy Partners And Interxon Energy Partners II |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |
Payments to acquire interest in joint venture | $ 7,000 |  |

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