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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenue (Details) - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
--- | --- | ---
Sep. 30, 2022 | Sep. 30, 2021 | Sep. 30, 2022 | Sep. 30, 2021
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | $ 16,722 | $ 3,333 | $ 25,146 | $ 10,583
Collaboration and licensing agreements |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | 14,561 | 22 | 14,561 | 389
Intrexon Energy Partners, LLC | Collaboration and licensing agreements |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | 3,768 | 0 | 3,768 | 0
Intrexon Energy Partners II, LLC | Collaboration and licensing agreements |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | 10,793 | 0 | 10,793 | 0
Castle Creek Biosciences, Inc. | Collaboration and licensing agreements |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | 0 | 18 | 0 | 371
Other | Collaboration and licensing agreements |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |
Total revenues | $ 0 | $ 4 | $ 0 | $ 18

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