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Condensed Consolidated Statements of Operations (Unaudited) - USD ($)
shares in Thousands, $ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2015
Sep. 30, 2014
Sep. 30, 2015
Sep. 30, 2014
Revenues:        
Collaboration revenue $ 829 $ 1,795 $ 2,230 $ 6,199
Grant revenue 359 414 608 2,546
Service and license revenue     527 10
Total revenues 1,188 2,209 3,365 8,755
Operating expenses:        
Research and development 24,105 28,072 66,011 64,942
General and administrative 5,524 4,083 15,481 12,325
Unoccupied facility expense   131   386
Total operating expenses 29,629 32,286 81,492 77,653
Loss from operations (28,441) (30,077) (78,127) (68,898)
Other income (expense):        
Interest income 33 42 78 162
Interest expense (62)   (572)  
Other income, net 17 216 360 300
Loss on extinguishment of debt (1,671)   (1,671)  
Net loss $ (30,124) $ (29,819) $ (79,932) $ (68,436)
Basic and diluted net loss per share $ (0.82) $ (1.13) $ (2.43) $ (2.60)
Weighted average shares used to compute basic and diluted net loss per share 36,532 26,291 32,880 26,288