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Condensed Consolidated Statements of Operations (Unaudited) - USD ($)
shares in Thousands, $ in Thousands
 3 Months Ended 6 Months Ended
Jun. 30, 2016
Jun. 30, 2015
Jun. 30, 2016
Jun. 30, 2015
Revenues:        
Collaboration revenue $ 1,683 $ 930 $ 2,578 $ 1,401
Grant revenue 88 101 127 249
Service and license revenue 876 519 884 527
Total revenues 2,647 1,550 3,589 2,177
Operating expenses:        
Research and development 22,750 19,686 42,817 41,906
General and administrative 9,151 5,098 17,320 9,957
Total operating expenses 31,901 24,784 60,137 51,863
Loss from operations (29,254) (23,234) (56,548) (49,686)
Other income (expense):        
Interest income 220 18 445 45
Interest expense   (263)   (510)
Other income (expense), net 48 (112) 94 343
Net loss $ (28,986) $ (23,591) $ (56,009) $ (49,808)
Basic and diluted net loss per share $ (0.75) $ (0.80) $ (1.46) $ (1.70)
Weighted average shares used to compute basic and diluted net loss per share 38,496 29,335 38,491 29,230