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Consolidated Statements of Stockholders' Equity - USD ($) shares in Thousands, $ in Thousands | Total | Common Stock | Preferred Stock | Additional Paid-in Capital | Accumulated Other Comprehensive Loss | Accumulated Deficit
--- | --- | --- | --- | --- | --- | ---
Beginning Balances at Dec. 31, 2014 | $ 100,482 | $ 26 |  | $ 695,058 | $ (1,669) | $ (592,933)
Beginning Balances (in shares) at Dec. 31, 2014 |  | 26,307 | 43 |  |  |
Conversion of Preferred Stock | 2 | $ 5 |  | (3) |  |
Conversion of Preferred Stock (in shares) |  | 4,343 | (43) |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | 531 |  |  | 531 |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  | 37 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 291 |  |  | 291 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 23 |  |  |  |
Issuance of common stock, net of issuance costs | 183,897 | $ 7 |  | 183,890 |  |
Issuance of common stock, net of issuance costs (in shares) |  | 7,353 |  |  |  |
Warrants exercised | 228 |  |  | 228 |  |
Warrants exercised (in shares) |  | 383 |  |  |  |
Stock compensation expense | 9,703 |  |  | 9,703 |  |
Total other comprehensive loss | (1,261) |  |  |  | (1,261) |
Net loss | (106,794) |  |  |  |  | (106,794)
Ending Balances at Dec. 31, 2015 | 187,079 | $ 38 |  | 889,698 | (2,930) | (699,727)
Ending Balances (in shares) at Dec. 31, 2015 |  | 38,446 |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | (83) | $ 1 |  | (84) |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  | 107 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 615 |  |  | 615 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 46 |  |  |  |
Stock compensation expense | 14,728 |  |  | 14,728 |  |
Total other comprehensive loss | (694) |  |  |  | (694) |
Net loss | (112,444) |  |  |  |  | (112,444)
Ending Balances at Dec. 31, 2016 | 89,201 | $ 39 |  | 904,957 | (3,624) | (812,171)
Ending Balances (in shares) at Dec. 31, 2016 |  | 38,599 |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | 1,613 |  |  | 1,613 |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  | 262 |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 293 |  |  | 293 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 84 |  |  |  |
Issuance of common stock, net of issuance costs | 185,936 | $ 23 |  | 185,913 |  |
Issuance of common stock, net of issuance costs (in shares) |  | 22,588 |  |  |  |
Stock compensation expense | 14,917 |  |  | 14,917 |  |
Total other comprehensive loss | 2,743 |  |  |  | 2,743 |
Net loss | (95,154) |  |  |  |  | (95,154)
Ending Balances at Dec. 31, 2017 | $ 199,549 | $ 62 |  | $ 1,107,693 | $ (881) | $ (907,325)
Ending Balances (in shares) at Dec. 31, 2017 |  | 61,533 |  |  |  |

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