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Condensed Consolidated Statements of Stockholders' Equity - USD ($) shares in Thousands, $ in Thousands | Total | Series B Convertible Preferred Stock | Common Stock | Preferred Stock | Preferred Stock Series B Convertible Preferred Stock | Additional Paid-in Capital | Additional Paid-in Capital Series B Convertible Preferred Stock | Accumulated Other Comprehensive (Loss) Income | Accumulated Deficit
--- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning Balances at Dec. 31, 2017 | $ 199,549 |  | $ 62 |  |  | $ 1,107,693 |  | $ (881) | $ (907,325)
Beginning Balances (in shares) at Dec. 31, 2017 |  |  | 61,533 |  |  |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | (529) |  | $ 1 |  |  | (530) |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  |  | 1,033 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 594 |  |  |  |  | 594 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 125 |  |  |  |  |  |
Stock compensation expense | 17,135 |  |  |  |  | 17,135 |  |  |
Total other comprehensive loss | (806) |  |  |  |  |  |  | (806) |
Net loss | (118,930) |  |  |  |  |  |  |  | (118,930)
Ending Balances at Sep. 30, 2018 | 97,013 |  | $ 63 |  |  | 1,124,892 |  | (1,687) | (1,026,255)
Ending Balances (in shares) at Sep. 30, 2018 |  |  | 62,691 |  |  |  |  |  |
Beginning Balances at Jun. 30, 2018 | 131,313 |  | $ 63 |  |  | 1,118,487 |  | (1,510) | (985,727)
Beginning Balances (in shares) at Jun. 30, 2018 |  |  | 62,608 |  |  |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | 20 |  |  |  |  | 20 |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  |  | 16 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 339 |  |  |  |  | 339 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 67 |  |  |  |  |  |
Stock compensation expense | 6,046 |  |  |  |  | 6,046 |  |  |
Total other comprehensive loss | (177) |  |  |  |  |  |  | (177) |
Net loss | (40,528) |  |  |  |  |  |  |  | (40,528)
Ending Balances at Sep. 30, 2018 | 97,013 |  | $ 63 |  |  | 1,124,892 |  | (1,687) | (1,026,255)
Ending Balances (in shares) at Sep. 30, 2018 |  |  | 62,691 |  |  |  |  |  |
Beginning Balances at Dec. 31, 2018 | 63,065 |  | $ 63 |  |  | 1,131,241 |  | (2,015) | (1,066,224)
Beginning Balances (in shares) at Dec. 31, 2018 |  |  | 62,862 |  |  |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net | 2 |  | $ 1 |  |  | 1 |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  |  | 969 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 565 |  |  |  |  | 565 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 122 |  |  |  |  |  |
Issuance of common stock and preferred stock, net of issuance costs | 60,113 | $ 12,061 | $ 20 |  |  | 60,093 | $ 12,061 |  |
Issuance of common stock and preferred stock, net of issuance costs (in shares) |  |  | 19,912 |  | 5 |  |  |  |
Stock compensation expense | 20,267 |  |  |  |  | 20,267 |  |  |
Total other comprehensive loss | (1,073) |  |  |  |  |  |  | (1,073) |
Net loss | (115,844) |  |  |  |  |  |  |  | (115,844)
Ending Balances at Sep. 30, 2019 | 39,156 |  | $ 84 |  |  | 1,224,228 |  | (3,088) | (1,182,068)
Ending Balances (in shares) at Sep. 30, 2019 |  |  | 83,865 | 5 |  |  |  |  |
Beginning Balances at Jun. 30, 2019 | 10,588 |  | $ 65 |  |  | 1,161,115 |  | (1,983) | (1,148,609)
Beginning Balances (in shares) at Jun. 30, 2019 |  |  | 65,155 |  |  |  |  |  |
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares) |  |  | 138 |  |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 158 |  |  |  |  | 158 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 47 |  |  |  |  |  |
Issuance of common stock and preferred stock, net of issuance costs | 46,165 | $ 12,061 | $ 19 |  |  | 46,146 | $ 12,061 |  |
Issuance of common stock and preferred stock, net of issuance costs (in shares) |  |  | 18,525 |  | 5 |  |  |  |
Stock compensation expense | 4,748 |  |  |  |  | 4,748 |  |  |
Total other comprehensive loss | (1,105) |  |  |  |  |  |  | (1,105) |
Net loss | (33,459) |  |  |  |  |  |  |  | (33,459)
Ending Balances at Sep. 30, 2019 | $ 39,156 |  | $ 84 |  |  | $ 1,224,228 |  | $ (3,088) | $ (1,182,068)
Ending Balances (in shares) at Sep. 30, 2019 |  |  | 83,865 | 5 |  |  |  |  |

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