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Condensed Consolidated Statements of Stockholders' Equity - USD ($)
shares in Thousands, $ in Thousands
Total
Common Stock
Preferred Stock
Additional Paid-in Capital
Accumulated Other Comprehensive (Loss) Income
Accumulated Deficit
Beginning Balances at Dec. 31, 2018 $ 63,065 $ 63   $ 1,131,241 $ (2,015) $ (1,066,224)
Beginning Balances (in shares) at Dec. 31, 2018   62,862        
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (18)     (18)    
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares)   740        
Issuance of common stock under Employee Stock Purchase Plan 407     407    
Issuance of common stock under Employee Stock Purchase Plan (in shares)   75        
Issuance of common stock, net of issuance costs, in conjunction with an At Market Sales Agreement 13,623 $ 2   13,621    
Issuance of common stock, net of issuance costs, in conjunction with an At Market Sales Agreement (in shares)   1,343        
Stock compensation expense 6,170     6,170    
Total other comprehensive loss (416)       (416)  
Net loss (39,672)         (39,672)
Ending Balances at Mar. 31, 2019 43,159 $ 65   1,151,421 (2,431) (1,105,896)
Ending Balances (in shares) at Mar. 31, 2019   65,020        
Beginning Balances at Dec. 31, 2019 8,290 $ 84   1,229,417 (2,387) (1,218,824)
Beginning Balances (in shares) at Dec. 31, 2019   83,871 5      
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (1) $ 1   (2)    
Issuance (withholding) of common stock upon exercise of stock options and restricted stock awards, net (in shares)   728        
Issuance of common stock under Employee Stock Purchase Plan 311     311    
Issuance of common stock under Employee Stock Purchase Plan (in shares)   91        
Issuance of common stock, net of issuance costs, in conjunction with an At Market Sales Agreement 14,229 $ 3   14,226    
Issuance of common stock, net of issuance costs, in conjunction with an At Market Sales Agreement (in shares)   2,909        
Stock compensation expense 1,778     1,778    
Total other comprehensive loss (188)       (188)  
Net loss (12,595)         (12,595)
Ending Balances at Mar. 31, 2020 $ 11,824 $ 88   $ 1,245,730 $ (2,575) $ (1,231,419)
Ending Balances (in shares) at Mar. 31, 2020   87,599 5