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Income Taxes - Schedule of Significant Components of Deferred Tax Assets and Liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
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Deferred tax assets: |  |
Net operating loss carryforwards | $ 564,712 | $ 456,284
Research and other credits | 139,996 | 132,994
Operating lease liabilities | 34,680 | 35,672
Stock-based compensation | 33,408 | 32,517
Liability related to the sale of future royalties | 23,757 | 32,737
Development derivative liability | 6,639 | 0
Other | 10,651 | 11,688
Deferred tax assets before valuation allowance | 813,843 | 701,892
Valuation allowance for deferred tax assets | (785,748) | (670,103)
Total deferred tax assets | 28,095 | 31,789
Deferred tax liabilities: |  |
Operating lease right-of-use assets | (27,204) | (29,707)
Other | (564) | (1,856)
Total deferred tax liabilities | (27,768) | (31,563)
Net deferred tax assets | $ 327 | $ 226

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