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Condensed Consolidated Statements of Stockholders’ Equity (Unaudited) - USD ($) $ in Thousands | Total | Follow-On Offering | At-The-Market Offering | License And Collaboration Agreement | Common Stock | Common Stock Follow-On Offering | Common Stock At-The-Market Offering | Common Stock License And Collaboration Agreement | Additional Paid-In Capital | Additional Paid-In Capital Follow-On Offering | Additional Paid-In Capital At-The-Market Offering | Additional Paid-In Capital License And Collaboration Agreement | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss)
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Beginning balance (in shares) at Dec. 31, 2022 | [1] |  |  |  |  | 3,292,163 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2022 |  | $ 76,106 |  |  |  | $ 5 | [1] |  |  |  |  |  | $ 606,918 |  |  |  | $ (530,644) | $ (173)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | 7,593 |  |  |  |  |  |  |  |  |  |  | 7,593 |  |  |  |  |
Issuance of common stock (in shares) | [1] |  |  |  |  | 560,253 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock |  | 18,096 |  |  |  | $ 1 | [1] |  |  |  |  |  | 18,095 |  |  |  |  |
Vesting of restricted stock awards (in shares) | [1] |  |  |  |  | 11,516 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) | [1] |  |  |  |  | (2,875) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (127) |  |  |  |  |  |  |  |  |  |  | (127) |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) | [1] |  |  |  |  | 2,970 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options |  | 101 |  |  |  |  |  |  |  |  |  |  | 101 |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 154 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 154
Net loss |  | (37,455) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (37,455) |
Ending balance (in shares) at Mar. 31, 2023 | [1] |  |  |  |  | 3,864,027 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Mar. 31, 2023 |  | 64,468 |  |  |  | $ 6 | [1] |  |  |  |  |  | 632,580 |  |  |  | (568,099) | (19)
Beginning balance (in shares) at Dec. 31, 2022 | [1] |  |  |  |  | 3,292,163 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2022 |  | 76,106 |  |  |  | $ 5 | [1] |  |  |  |  |  | 606,918 |  |  |  | (530,644) | (173)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 173 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Net loss |  | (96,399) |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Ending balance (in shares) at Sep. 30, 2023 | [1] |  |  |  |  | 8,569,822 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Sep. 30, 2023 |  | 86,756 |  |  |  | $ 13 | [1] |  |  |  |  |  | 713,786 |  |  |  | (627,043) | 0
Beginning balance (in shares) at Mar. 31, 2023 | [1] |  |  |  |  | 3,864,027 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Mar. 31, 2023 |  | 64,468 |  |  |  | $ 6 | [1] |  |  |  |  |  | 632,580 |  |  |  | (568,099) | (19)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | 5,775 |  |  |  |  |  |  |  |  |  |  | 5,775 |  |  |  |  |
Issuance of common stock (in shares) | [1] |  |  |  |  |  |  | 4,296,646 |  | 392,541 |  |  |  |  |  |  |  |
Issuance of common stock |  |  | $ 63,439 | $ 6,032 |  |  |  | $ 6 | [1] | $ 1 | [1] |  |  | $ 63,433 | $ 6,031 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | [1] |  |  |  |  | 15,663 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan |  | 208 |  |  |  |  |  |  |  |  |  |  | 208 |  |  |  |  |
Vesting of restricted stock awards (in shares) | [1] |  |  |  |  | 720 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) | [1] |  |  |  |  | (249) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (4) |  |  |  |  |  |  |  |  |  |  | (4) |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 19 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 19
Net loss |  | (34,312) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (34,312) |
Ending balance (in shares) at Jun. 30, 2023 | [1] |  |  |  |  | 8,569,348 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Jun. 30, 2023 |  | 105,625 |  |  |  | $ 13 | [1] |  |  |  |  |  | 708,023 |  |  |  | (602,411) | 0
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | 5,763 |  |  |  |  |  |  |  |  |  |  | 5,763 |  |  |  |  |
Vesting of restricted stock awards (in shares) | [1] |  |  |  |  | 233 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) | [1] |  |  |  |  | (70) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (1) |  |  |  |  |  |  |  |  |  |  | (1) |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) | [1] |  |  |  |  | 311 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options |  | 1 |  |  |  |  |  |  |  |  |  |  | 1 |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Net loss |  | (24,632) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (24,632) |
Ending balance (in shares) at Sep. 30, 2023 | [1] |  |  |  |  | 8,569,822 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Sep. 30, 2023 |  | $ 86,756 |  |  |  | $ 13 | [1] |  |  |  |  |  | 713,786 |  |  |  | (627,043) | 0
Beginning balance (in shares) at Dec. 31, 2023 |  | 8,791,877 |  |  |  | 8,791,877 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2023 |  | $ 69,669 |  |  |  | $ 13 |  |  |  |  |  |  | 723,577 |  |  |  | (653,921) | 0
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | 14,475 |  |  |  |  |  |  |  |  |  |  | 14,475 |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  |  |  |  | 3,802,025 |  | 209,852 |  | 443,253 |  |  |  |  |  |
Issuance of common stock |  |  | 161,649 | $ 6,169 | $ 17,265 |  |  |  |  |  |  |  |  | 161,649 | $ 6,169 | $ 17,265 |  |
Vesting of restricted stock awards (in shares) |  |  |  |  |  | 11,095 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) |  |  |  |  |  | (3,689) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (137) |  |  |  |  |  |  |  |  |  |  | (137) |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) |  |  |  |  |  | 3,634 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options |  | 143 |  |  |  |  |  |  |  |  |  |  | 143 |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 3 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 3
Net loss |  | (39,553) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (39,553) |
Ending balance (in shares) at Mar. 31, 2024 |  |  |  |  |  | 13,258,047 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Mar. 31, 2024 |  | $ 229,683 |  |  |  | $ 13 |  |  |  |  |  |  | 923,141 |  |  |  | (693,474) | 3
Beginning balance (in shares) at Dec. 31, 2023 |  | 8,791,877 |  |  |  | 8,791,877 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Dec. 31, 2023 |  | $ 69,669 |  |  |  | $ 13 |  |  |  |  |  |  | 723,577 |  |  |  | (653,921) | 0
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) |  | 8,753 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | $ 1,331 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Net loss |  | $ (124,140) |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Ending balance (in shares) at Sep. 30, 2024 |  | 17,785,697 |  |  |  | 17,785,697 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Sep. 30, 2024 |  | $ 382,666 |  |  |  | $ 14 |  |  |  |  |  |  | 1,159,382 |  |  |  | (778,061) | 1,331
Beginning balance (in shares) at Mar. 31, 2024 |  |  |  |  |  | 13,258,047 |  |  |  |  |  |  |  |  |  |  |  |
Beginning balance at Mar. 31, 2024 |  | 229,683 |  |  |  | $ 13 |  |  |  |  |  |  | 923,141 |  |  |  | (693,474) | 3
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | $ 5,878 |  |  |  |  |  |  |  |  |  |  | 5,878 |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  |  |  |  | 3,849,558 |  |  |  |  |  |  |  |  |  |
Issuance of common stock |  |  | 215,988 |  |  |  |  | $ 1 |  |  |  |  |  | 215,987 |  |  |  |
Issuance of common stock from exercise of pre-funded warrants (in shares) |  | 622,123 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  |  |  |  | 25,146 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock under employee stock purchase plan |  | $ 311 |  |  |  |  |  |  |  |  |  |  | 311 |  |  |  |  |
Vesting of restricted stock awards (in shares) |  |  |  |  |  | 726 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) |  |  |  |  |  | (242) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (13) |  |  |  |  |  |  |  |  |  |  | (13) |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) |  |  |  |  |  | 148 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options |  | 4 |  |  |  |  |  |  |  |  |  |  | 4 |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | (74) |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (74)
Net loss |  | (32,677) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (32,677) |
Ending balance (in shares) at Jun. 30, 2024 |  |  |  |  |  | 17,755,506 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Jun. 30, 2024 |  | 419,100 |  |  |  | $ 14 |  |  |  |  |  |  | 1,145,308 |  |  |  | (726,151) | (71)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  | 12,432 |  |  |  |  |  |  |  |  |  |  | 12,432 |  |  |  |  |
Issuance of common stock (in shares) |  |  |  |  |  |  |  | 25,189 |  |  |  |  |  |  |  |  |  |
Issuance of common stock |  |  | $ 1,442 |  |  |  |  |  |  |  |  |  |  | $ 1,442 |  |  |  |
Vesting of restricted stock awards (in shares) |  |  |  |  |  | 51 |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units (in shares) |  |  |  |  |  | (20) |  |  |  |  |  |  |  |  |  |  |  |
Shares withheld for taxes for vesting of restricted stock units |  | (1) |  |  |  |  |  |  |  |  |  |  | (1) |  |  |  |  |
Issuance of common stock upon exercise of stock options (in shares) |  |  |  |  |  | 4,971 |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options |  | 201 |  |  |  |  |  |  |  |  |  |  | 201 |  |  |  |  |
Change in unrealized gain on marketable securities, net of tax |  | 1,402 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 1,402
Net loss |  | $ (51,910) |  |  |  |  |  |  |  |  |  |  |  |  |  |  | (51,910) |
Ending balance (in shares) at Sep. 30, 2024 |  | 17,785,697 |  |  |  | 17,785,697 |  |  |  |  |  |  |  |  |  |  |  |
Ending balance at Sep. 30, 2024 |  | $ 382,666 |  |  |  | $ 14 |  |  |  |  |  |  | $ 1,159,382 |  |  |  | $ (778,061) | $ 1,331
[1] | Results have been retroactively adjusted to reflect the 1-for-15 reverse stock split effected on November 28, 2023. See Note 2 for details.

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