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STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Deficit | Accumulated Other Comprehensive Gain (Loss)
--- | --- | --- | --- | --- | ---
Balances at Dec. 31, 2018 | $ 177,733 | $ 186 | $ 1,189,194 | $ (1,011,464) | $ (183)
Balances (in shares) at Dec. 31, 2018 |  | 186,392,682 |  |  |
Increase (Decrease) in Stockholders' Equity |  |  |  |  |
Net loss | (68,548) |  |  | (68,548) |
Other comprehensive income (loss) | 315 |  |  |  | 315
Issuance of common stock in connection with at market offering, net of issuance costs | 19,295 | $ 14 | 19,281 |  |
Issuance of common stock in connection with at market offering, net of issuance costs (in shares) |  | 13,214,867 |  |  |
Stock-based compensation related to issuance of common stock and options in exchange for services | 68 |  | 68 |  |
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares) |  | 29,150 |  |  |
Issuances of common stock under equity plans | 204 |  | 204 |  |
Issuances of common stock under equity plans (in shares) |  | 177,882 |  |  |
Stock-based compensation for equity-based awards to employees and directors | 6,079 |  | 6,079 |  |
401(k) contribution | 9 |  | 9 |  |
Balances at Dec. 31, 2019 | 135,155 | $ 200 | 1,214,835 | (1,080,012) | 132
Balances (in shares) at Dec. 31, 2019 |  | 199,814,581 |  |  |
Increase (Decrease) in Stockholders' Equity |  |  |  |  |
Net loss | (75,617) |  |  | (75,617) |
Other comprehensive income (loss) | (54) |  |  |  | (54)
Issuance of common stock, pre-funded warrant and warrants to purchase common stock in public offering, net of issuance costs | 140,184 | $ 107 | 140,077 |  |
Issuance of common stock, pre-funded warrant and warrants to purchase common stock in public offering, net of issuance costs (in shares) |  | 107,049,375 |  |  |
Issuance of common stock in connection with at market offering, net of issuance costs | 4,075 | $ 3 | 4,072 |  |
Issuance of common stock in connection with at market offering, net of issuance costs (in shares) |  | 3,496,616 |  |  |
Issuance of common stock in connection exercise of warrants | 16 |  | 16 |  |
Issuance of common stock in connection exercise of warrants (in shares) |  | 12,500 |  |  |
Stock-based compensation related to issuance of common stock and options in exchange for services | 85 |  | 85 |  |
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares) |  | 17,986 |  |  |
Issuances of common stock under equity plans | 208 |  | 208 |  |
Issuances of common stock under equity plans (in shares) |  | 175,795 |  |  |
Stock-based compensation for equity-based awards to employees and directors | 6,895 |  | 6,895 |  |
Balances at Dec. 31, 2020 | $ 210,947 | $ 310 | 1,366,188 | (1,155,629) | 78
Balances (in shares) at Dec. 31, 2020 | 310,566,853 | 310,566,853 |  |  |
Increase (Decrease) in Stockholders' Equity |  |  |  |  |
Net loss | $ (116,112) |  |  | (116,112) |
Other comprehensive income (loss) | (251) |  |  |  | (251)
Issuance of common stock in connection with at market offering, net of issuance costs | 20,385 | $ 11 | 20,374 |  |
Issuance of common stock in connection with at market offering, net of issuance costs (in shares) |  | 10,571,556 |  |  |
Issuance of common stock in connection exercise of warrants | 2,479 | $ 2 | 2,477 |  |
Issuance of common stock in connection exercise of warrants (in shares) |  | 1,906,341 |  |  |
Stock-based compensation related to issuance of common stock and options in exchange for services | 91 |  | 91 |  |
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares) |  | 20,783 |  |  |
Issuances of common stock under equity plans | 797 | $ 1 | 796 |  |
Issuances of common stock under equity plans (in shares) |  | 666,058 |  |  |
Stock-based compensation for equity-based awards to employees and directors | 8,080 |  | 8,080 |  |
Balances at Dec. 31, 2021 | $ 126,416 | $ 324 | $ 1,398,006 | $ (1,271,741) | $ (173)
Balances (in shares) at Dec. 31, 2021 | 323,731,591 | 323,731,591 |  |  |

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