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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
Total
Common Stock
Additional Paid-In Capital
Accumulated Deficit
Accumulated Other Comprehensive Gain (Loss)
Balances at Dec. 31, 2020 $ 210,947,000 $ 310,000 $ 1,366,188,000 $ (1,155,629,000) $ 78,000
Balances (in shares) at Dec. 31, 2020   310,566,853      
Increase (Decrease) in Stockholders' Equity          
Net loss (27,824,000)     (27,824,000)  
Other comprehensive income (loss) (43,000)       (43,000)
Issuance of common stock in connection with at market offering, net of issuance costs 16,234,000 $ 8,000 16,226,000    
Issuance of common stock in connection with at market offering, net of issuance costs (in shares)   7,948,505      
Stock-based compensation related to issuance of common stock and options in exchange for services 25,000   25,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   4,549      
Issuance of common stock in connection with exercise of warrants 12,000   12,000    
Issuance of common stock in connection exercise of warrants (in shares)   8,869      
Issuances of common stock under equity plans 17,000   17,000    
Issuances of common stock under equity plans (in shares)   16,232      
Stock-based compensation for equity-based awards to employees and directors 1,794,000   1,794,000    
Balances at Mar. 31, 2021 201,162,000 $ 318,000 1,384,262,000 (1,183,453,000) 35,000
Balances (in shares) at Mar. 31, 2021   318,545,008      
Balances at Dec. 31, 2020 210,947,000 $ 310,000 1,366,188,000 (1,155,629,000) 78,000
Balances (in shares) at Dec. 31, 2020   310,566,853      
Increase (Decrease) in Stockholders' Equity          
Net loss (84,095,000)        
Comprehensive loss (84,157,000)        
Balances at Sep. 30, 2021 151,553,000 $ 321,000 1,390,940,000 (1,239,724,000) 16,000
Balances (in shares) at Sep. 30, 2021   320,604,441      
Balances at Mar. 31, 2021 201,162,000 $ 318,000 1,384,262,000 (1,183,453,000) 35,000
Balances (in shares) at Mar. 31, 2021   318,545,008      
Increase (Decrease) in Stockholders' Equity          
Net loss (29,574,000)     (29,574,000)  
Other comprehensive income (loss) (32,000)       (32,000)
Stock-based compensation related to issuance of common stock and options in exchange for services 23,000   23,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   5,097      
Issuance of common stock in connection with exercise of warrants 2,467,000 $ 2,000 2,465,000    
Issuance of common stock in connection exercise of warrants (in shares)   1,897,472      
Issuances of common stock under equity plans 187,000 $ 1,000 186,000    
Issuances of common stock under equity plans (in shares)   151,618      
Stock-based compensation for equity-based awards to employees and directors 1,973,000   1,973,000    
Balances at Jun. 30, 2021 176,206,000 $ 321,000 1,388,909,000 (1,213,027,000) 3,000
Balances (in shares) at Jun. 30, 2021   320,599,195      
Increase (Decrease) in Stockholders' Equity          
Net loss (26,697,000)     (26,697,000)  
Other comprehensive income (loss) 13,000       13,000
Comprehensive loss (26,684,000)        
Stock-based compensation related to issuance of common stock and options in exchange for services 23,000   23,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   5,246      
Stock-based compensation for equity-based awards to employees and directors 2,008,000   2,008,000    
Balances at Sep. 30, 2021 151,553,000 $ 321,000 1,390,940,000 (1,239,724,000) 16,000
Balances (in shares) at Sep. 30, 2021   320,604,441      
Balances at Dec. 31, 2021 126,416,000 $ 324,000 1,398,006,000 (1,271,741,000) (173,000)
Balances (in shares) at Dec. 31, 2021   323,731,591      
Increase (Decrease) in Stockholders' Equity          
Net loss (30,098,000)     (30,098,000)  
Other comprehensive income (loss) (538,000)       (538,000)
Stock-based compensation related to issuance of common stock and options in exchange for services 15,000   15,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   5,284      
Stock-based compensation for equity-based awards to employees and directors 1,692,000   1,692,000    
Balances at Mar. 31, 2022 97,487,000 $ 324,000 1,399,713,000 (1,301,839,000) (711,000)
Balances (in shares) at Mar. 31, 2022   323,736,875      
Balances at Dec. 31, 2021 126,416,000 $ 324,000 1,398,006,000 (1,271,741,000) (173,000)
Balances (in shares) at Dec. 31, 2021   323,731,591      
Increase (Decrease) in Stockholders' Equity          
Net loss (99,266,000)        
Comprehensive loss (99,713,000)        
Balances at Sep. 30, 2022 108,172,000 $ 382,000 1,479,417,000 (1,371,007,000) (620,000)
Balances (in shares) at Sep. 30, 2022   381,214,326      
Balances at Mar. 31, 2022 97,487,000 $ 324,000 1,399,713,000 (1,301,839,000) (711,000)
Balances (in shares) at Mar. 31, 2022   323,736,875      
Increase (Decrease) in Stockholders' Equity          
Net loss (28,117,000)     (28,117,000)  
Other comprehensive income (loss) (71,000)       (71,000)
Issuance of common stock, pre-funded warrant and warrants to purchase common stock in public offering, net of issuance costs 69,916,000 $ 53,000 69,863,000    
Issuance of common stock, pre-funded warrant and warrants to purchase common stock in public offering, net of issuance costs, (in shares)   53,333,334      
Stock-based compensation related to issuance of common stock and options in exchange for services 13,000   13,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   4,637      
Issuances of common stock under equity plans 761,000 $ 1,000 760,000    
Issuances of common stock under equity plans (in shares)   589,913      
Stock-based compensation for equity-based awards to employees and directors 2,058,000   2,058,000    
Balances at Jun. 30, 2022 142,047,000 $ 378,000 1,472,407,000 (1,329,956,000) (782,000)
Balances (in shares) at Jun. 30, 2022   377,664,759      
Increase (Decrease) in Stockholders' Equity          
Net loss (41,051,000)     (41,051,000)  
Other comprehensive income (loss) 162,000       162,000
Comprehensive loss (40,889,000)        
Stock-based compensation related to issuance of common stock and options in exchange for services 112,000   112,000    
Stock-based compensation related to issuance of common stock and options in exchange for services (in shares)   3,071      
Issuance of common stock in connection with exercise of warrants 3,638,000 $ 3,000 3,635,000    
Issuance of common stock in connection exercise of warrants (in shares)   2,798,003      
Issuances of common stock under equity plans 1,201,000 $ 1,000 1,200,000    
Issuances of common stock under equity plans (in shares)   748,493      
Stock-based compensation for equity-based awards to employees and directors 2,063,000   2,063,000    
Balances at Sep. 30, 2022 $ 108,172,000 $ 382,000 $ 1,479,417,000 $ (1,371,007,000) $ (620,000)
Balances (in shares) at Sep. 30, 2022   381,214,326