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Stock Based Compensation (Tables)

3 Months Ended

Mar. 31, 2019

Disclosure Of Compensation Related Costs Sharebased Payments [Abstract]

Summary of Option Activity Under Company's Equity Incentive Plans

The following table summarizes stock option activity under the Company’s equity incentive plans and related information:

		Number of Shares Underlying Outstanding Options				Weighted- Average Exercise Price				Weighted- Average Remaining Contractual Term (Years)
Outstanding — December 31, 2018			1,361,977			$	12.04				8.71
Options granted			309,300			$	8.12
Options exercised			(12,794	)		$	4.60
Options forfeited			(39,971	)		$	7.60
Options Expired			(5,641	)		$	91.79
Outstanding — March 31, 2019			1,612,871			$	11.17				8.77
Exercisable — March 31, 2019			468,193			$	16.19				8.03
Vested and expected to vest — March 31, 2019			1,612,871			$	11.17				8.77

Summary of Weighted Average Valuation Assumptions Used to Estimate Fair Value of Employee Stock Options

The fair value of employee stock options was estimated using the following weighted-average assumptions for the three months ended March 31, 2019 and 2018:

		Three Months Ended March 31,
		2019				2018
Employee Stock Options:
Risk-free interest rate			2.56	%			2.40	%
Expected term (in years)			6.03				6.01
Dividend yield			—				—
Volatility			82.91	%			109.32	%
Weighted-average fair value of stock options granted		$	5.80			$	12.34

Summary of Stock-Based Compensation Recognized

Total stock-based compensation recognized was as follows (in thousands):

		Three Months Ended March 31,
		2019				2018
Research and development		$	238			$	115
General and administrative(1)			591				491
Total stock-based compensation		$	829			$	606

(1) 2018 includes $0.1 million in modification stock-based compensation expense related to a Board member’s departure.

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Tabular disclosure of the allocation of equity-based compensation costs to a given line item on the balance sheet and income statement for the period. This may include the reporting line for the costs and the amount capitalized and expensed.

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Tabular disclosure for stock option plans. Includes, but is not limited to, outstanding awards at beginning and end of year, grants, exercises, forfeitures, and weighted-average grant date fair value.

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Tabular disclosure of the significant assumptions used during the year to estimate the fair value of stock options, including, but not limited to: (a) expected term of share options and similar instruments, (b) expected volatility of the entity's shares, (c) expected dividends, (d) risk-free rate(s), and (e) discount for post-vesting restrictions.

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