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Collaborations - Additional Information (Detail) - USD ($) |  | 1 Months Ended | 3 Months Ended
--- | --- | --- | ---
May 08, 2020 | Jan. 31, 2020 | Dec. 31, 2018 | Mar. 31, 2022 | Mar. 31, 2021
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
License and collaboration revenue |  |  |  | $ 794,000 | $ 1,467,000
Research and development |  |  |  | 9,703,000 | 17,013,000
Deferred revenue, recognized |  |  |  | 200,000 |
Collaboration |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
License and collaboration revenue |  |  |  | 794,000 | 1,467,000
Collaboration | Reimbursable Out-of-Pocket and Personnel Costs Incurred Related to Research Services |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
License and collaboration revenue |  |  |  | 800,000 | 1,500,000
Mosaic | Maximum |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Milestone payment related to regulatory and clinical development events | $ 4,000,000.0 |  |  |  |
ISU Abxis | Amended and Restated License Agreement |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Regulatory and development milestone payment |  |  | $ 2,500,000 | 0 |
ISU Abxis | Maximum | Amended and Restated License Agreement |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Milestone payment |  |  | 19,500,000 | 0 |
Commercial milestone payments |  |  | $ 17,000,000.0 | 0 |
Biogen |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
License and collaboration revenue |  |  |  | 0 | 0
Biogen | Collaboration |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Research and development |  |  |  | $ 700,000 | $ 1,400,000
Biogen | Exclusive License |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Up-front payments received |  | $ 15,000,000.0 |  |  |
Biogen | Maximum | Exclusive License |  |  |  |  |
Revenue Recognition Multiple Deliverable Arrangements [Line Items] |  |  |  |  |
Development and sales milestone payments |  | $ 340,000,000.0 |  |  |

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- Definition The aggregate costs incurred (1) in a planned search or critical investigation aimed at discovery of new knowledge with the hope that such knowledge will be useful in developing a new product or service, a new process or technique, or in bringing about a significant improvement to an existing product or process; or (2) to translate research findings or other knowledge into a plan or design for a new product or process or for a significant improvement to an existing product or process whether intended for sale or the entity's use, during the reporting period charged to research and development projects, including the costs of developing computer software up to the point in time of achieving technological feasibility, and costs allocated in accounting for a business combination to in-process projects deemed to have no alternative future use. + References Reference 1: http://www.xbrl.org/2009/role/commonPracticeRef -Publisher FASB -Name Accounting Standards Codification -Topic 912 -SubTopic 730 -Section 25 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=6472174&loc=d3e58812-109433 Reference 2: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 985 -SubTopic 20 -Section 50 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=6501960&loc=d3e128462-111756 Reference 3: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 730 -SubTopic 10 -Section 50 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=6420194&loc=d3e21568-108373 + Details
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